Research Article | | Peer-Reviewed

New Ranks and New Combinations in Selected Families and Genera

Published in Plant (Volume 13, Issue 2)
Received: 13 May 2025     Accepted: 26 May 2025     Published: 20 June 2025
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Abstract

Recent advancements in molecular phylogenetics have significantly reshaped infrageneric classifications across vascular plant genera, yet nomenclature within these ranks remains understudied. This article addresses critical gaps in the taxonomy of selected plant families by introducing nomenclatural novelties aligned with modern systematic frameworks. The study is structured into three interrelated sections. First, updated infrageneric classifications are proposed for six genera, i. e., Calanthe, Docynia, Epimedium, Eriolobus, Galanthus, and Nymphaea, integrating morphological and molecular evidence to resolve longstanding taxonomic ambiguities. For instance, formerly segregated genera Cephalantheropsis, Gastrorchis, and Phaius is herein newly treated as subgenera under the broadly circumscribed Calanthe, and Paraphaius is treated as a synonym of Calanthe subgen. Hecabe; while Docynia is retained as distinct from Malus due to its unique ecological and evolutionary adaptations. The second section validates 373 new nothoranks across 58 genera from 29 families of pteridophytes and spermatophytes: Amaranthaceae, Amaryllidaceae, Anemiaceae, Asparagaceae, Asphodelaceae, Asteraceae, Athyriaceae, Berberidaceae, Cistaceae, Crassulaceae, Dryopteridaceae, Heliconiaceae, Hydrangeaceae, Hypericaceae, Iridaceae, Lamiaceae, Lentibulariaceae, Marattiaceae, Musaceae, Nymphaeaceae, Orchidaceae, Papaveraceae, Plantaginaceae, Plumbaginaceae, Ranunculaceae, Rosaceae, Saxifragaceae, Verbenaceae and Violaceae. These include novel nothosections, nothosubsections, nothoseries, and nothosubgenera to accommodate hybrids previously lacking suitable placement. The third section establishes three new nothogenera and 28 new combinations across nine families, namely Apocynaceae, Athyriaceae, Convolvulaceae, Lamiaceae, Orchidaceae, Poaceae, Rosaceae, Saxifragaceae and Thelypteridaceae, resolving taxonomic inconsistencies arising from recent phylogenetic insights.

Published in Plant (Volume 13, Issue 2)
DOI 10.11648/j.plant.20251302.14
Page(s) 76-99
Creative Commons

This is an Open Access article, distributed under the terms of the Creative Commons Attribution 4.0 International License (http://creativecommons.org/licenses/by/4.0/), which permits unrestricted use, distribution and reproduction in any medium or format, provided the original work is properly cited.

Copyright

Copyright © The Author(s), 2025. Published by Science Publishing Group

Keywords

Infrageneric Classification, New Combination, Nothogenus, Nothorank

1. Introduction
At familial and infrafamilial level, the taxonomic framework is rather steady with combination of more extensive and integrate morphological and molecular studies, while most infrageneric classification are either reexamined or recircumscribed in two decades phylogenetically. Contrast to the creation and publishment of new nothogenera and nothospecies, the attention on infrageneric hybrid names is still not enough in most families of vascular plants. From 1973-2021, there are 256 published infrafamilial nothoranks (24 nothosubgenera, 185 nothosections, 20 nothosubsections and 23 nothoseries) for 44 (notho) genera from 22 families, plus two nothosubtribes in Cactaceae. The current article aims to: (1) present updated infrageneric classificatory scheme for several genera with evidences from recent advances regarding their taxonomy; (2) integrate appropriate required infrageneric nothoranks to accommodate certain named nothospecies that are umcommonly provided in extant families and genera, according to the most recent or reasonable infrageneric classification for the involved genera; (3) mandate new nothogenera and combinations to meet recent taxonomic changes.
2. Sources and Methods
2.1. Literature Survey and Classification
A dataset of recorded hybrid names up to 1 May 2025 was compiled from online databases as the original material of current work. Protologues are checked in an effort to ensure accuracy of the hybrid formula of involved taxa, but as there are too many to cite, all of them can be found online. At the family level, the modern classification systems - APG IV for flowering plants, and PPG I for ferns - are adopted. Infrageneric taxonomic framework and species concepts adopted here following particular phylogenetic studies and monographic works, are otherwise explained. Revised infrageneric classification proposed for selected genera herein are represent of current knowledge and understanding utilizing morphology and monophyletics. Further details are provided to make precise notes.
2.2. Nomenclature
Shenzhen Code of ICN has been followed in making necessary nomenclatural decisions. Unless more recent information is available, author attributions of scientific plant names cited follows IPNI , albeit in the notation required by the journal. In addition to their use in direct citations of text(s), in accordance with the notation used by ICN (: 205, ‘Index to scientific names’), double quotation marks [“ ”] are placed around designations or orthographic varieties to indicate that they are not ‘names’ in the sense of ICN (: Art. 6.3).
2.3. Formatting
Unless more recent information is available, author attribution and original publication of nothospecific names are in the notation of IPNI. Herbarium codes follow Thiers. In the following treatments, all names are arranged in alphabetical order to ease the use of the checklist. To achieve precise writing, only selected references are provided for crucial taxonomic information, a brief note is provided when necessary to make clarifications.
3. Nomenclatural Treatment
3.1. Updated Infrageneric Classification for Six Genera
Amaryllidaceae
Galanthus L.
Remark: The morphology-based infrageneric classification of Galanthus is not fully consistent with phylogenetic findings. Based on phylogenetic results , Galanthus can be divided into seven series, which are corresponding to its seven well-supported clades: ser. Latifolii Kem.-Nath. (Platyphyllus clade), ser. Trojanenses Z. H. Feng (Trojanus clade), ser. Ikarienses Z. H. Feng (Ikariae clade), ser. Elwesiani Kem.-Nath. ex A. P. Khokhr. (Elwesii clade), ser. Galanthus (Nivalis clade), ser. Woronowii Kem.-Nath. (Woronowii clade) and ser. Alpini Z. H. Feng (Alpinus clade).
Galanthus ser. Latifolii Kem.-Nath. (1947).
Galanthus ser. Trojanenses Z. H. Feng, ser. nov. Type: Galanthus trojanus A. P. Davis & Özhatay.
Diagnosis: Bulbs ± ovoid or spherical; vernation applanate; leaves linear, sometimes slightly oblanceolate, medium to dark green; inner perianth segments with an apical notch, and a single green mark at the apex, or sometimes with two eye-like marks at the base. Monotypic.
Galanthus ser. Ikarienses Z. H. Feng, ser. nov. Type: Galanthus ikariae Baker.
Diagnosis: Bulbs ± ovoid or spherical; vernation supervoloute; leaves broad-linear to narrowly oblanceolate, light to medium green; inner perianth segments with an apical notch, and a single bold green mark (circa half the size of the segment) at the apex. Monotypic.
Galanthus ser. Elwesiani Kem.-Nath. ex A. P. Khokhr. (1966).
Galanthus ser. Galanthus.
Galanthus ser. Woronowii Kem.-Nath. (1947).
Galanthus ser. Alpini Z. H. Feng, ser. nov. Type: Galanthus alpinus Sosnowsky.
Diagnosis: Bulbs ± ovoid or spherical; vernation supervolute or applanate (only G. angustifolius); leaves linear (G. angustifolius, G. koenenianus) to narrowly oblanceolate, glaucous or greenglaucescent (G. transcaucasicus); inner perianth segments with an apical notch, and a single green mark at the apex, sometimes with a faint yellowish or green mark at the base (only G. koenenianus). Four species: G. alpinus, G. angustifolius, G. koenenianus and G. transcaucasicus.
Berberidaceae
Epimedium L.
Remark: The serial classification of Epimedium sect. Diphyllon is controversial by different authors using floral spur, number of cauline leaf, rhizome or geographical distribution. According to the subsequent molecular findings, Chinese sect. Diphyllon was divided into five well-supported clades related to flower morphology except that five species were either isolated or formed a general polytomy.
Epimedium subgen. Rhizophyllum (Fisch. & C. A. Mey.) Stearn (2002).
Epimedium subgen. Epimedium
Epimedium sect. Diphyllon (Kom.) Stearn (2002).
Epimedium ser. Calcar (S. Z. He) Z. H. Feng, stat. nov. — Basionym: Epimedium sect. Calcar S. Z. He, Genus Epimedium China: 28. 2014.
Epimedium ser. Davidianae Stearn (2002).
Epimedium ser. Diphyllon Kom. (1908).
Epimedium ser. Dolichocera Stearn (1938).
Epimedium ser. Fargesii Z. H. Feng, ser. nov. Type: Epimedium fargesii Franch.
Diagnosis: Flowers small but slightly larger than those of Epimedium ser. Diphyllon, stamens conspicuously protruded, 8-10 mm long. Four species: E. dewuense, E. dolichostemon, E. fargesii and E. qingchengshanense.
Epimedium ser. Leptorrhizi Z. H. Feng, ser. nov. Type: Epimedium leptorrhizum Stearn.
Diagnosis: Rhizome long-creeping, 1-2 mm thick, with internodes sometimes as much as 20 cm long. Petioles furnished with spreading reddish hairs; lateral leaflet often pruinose abaxially. Monotypic.
Epimedium ser. Stellulati Z. H. Feng, ser. nov. Type: Epimedium stellulatum Stearn.
Diagnosis: Inflorescence compound, loose, circa 15-20 cm long, pedicels with numerous glandular hairs. Flowers small, Petals much shorter than inner sepals, with blunt nectariferous spur. Monotypic.
Epimedium sect. Epimedium
Epimedium sect. Macroceras C. Morren & Decne. (1834).
Epimedium sect. Polyphyllon (Kom.) Stearn (2002).
Nymphaeaceae
Nymphaea L.
Remark: Tradionally, Nymphaea is divided into five subgenera based on morphological traits, such as filament appearance, sterile appendage and carpel walls free or fused. Jacobs, in 2007, added a new subgenus Confluentes, characterized by flowers with a distinct gap between petals and stamens, leaf margins toothed and seeds rather large. According to the subsequent molecular findings , it is warrant to reduce Confluentes as a section within subgen. Anecphya s. lat.
Nymphaea subgen. Anecphya (Casp.) Conard (1905).
Nymphaea sect. Confluentes (S. W. L. Jacobs) Bing Liu ex Zi Jun Li & Z. H. Feng, stat. nov. — Basionym: Nymphaea subgen. Confluentes S. W. L. Jacobs, Fl. Australia 2: 458. 2007.
Nymphaea sect. Anecphya (Casp.) Bing Liu ex Zi Jun Li & Z. H. Feng, stat. nov. — Basionym: Nymphaea subsect. Anecphya Casp., Ann. Mus. Bot. Lugduno-Batavi 2: 247. 1866.
Nymphaea subgen. Brachyceras (Casp.) Conard (1905).
Nymphaea subgen. Nymphaea
Nymphaea sect. Xanthantha (Casp.) Wiersema (1996).
Nymphaea sect. Nymphaea
Nymphaea subgen. Lotos (DC.) Conard (1905).
Nymphaea subgen. Hydrocallis (Planch.) Conard (1905).
Orchidaceae
Calanthe R. Br.
Remark: In Calanthe, seven major subclades are found by molecular phylogenetic analyses covering the major infrageneric groups in Calanthe and Phaius plus representative species of allied genera. Among which, subclade (6) was elevated to generic level as Paraphaius. However, Chase et al. sought a broader delimitation of Calanthe to include the species of Cephalantheropsis, Gastrorchis and Phaius because “segregate new genera from both Phaius and Calanthe … results in generic concepts for which there are no reasonable morphological differences”, and since Calanthe is conserved against Phaius , a broad sense of Calanthe is here adopted with an updated infrageneric classification.
Calanthe subgen. Calanthe
Calanthe sect. Aceratochilus Schltr. (1912).
Calanthe sect. Alpinocalanthe J. W. Zhai, Z. J. Liu & F. W. Xing (2014).
Calanthe sect. Ghiesbreghtia (A. Rich. & Galeotti) Schltr. (1912).
Calanthe sect. Monophylla D. A. Clayton & P. J. Cribb (2013).
Calanthe sect. Puberula J. W. Zhai, Z. J. Liu & F. W. Xing (2014).
Calanthe sect. Tricarinata J. W. Zhai, Z. J. Liu & F. W. Xing (2014).
Calanthe sect. Verna Kakadzu (1994).
Calanthe subgen. Cephalantheropsis (Guillaumin) Z. H. Feng, comb. & stat. nov. — Basionym: Cephalantheropsis Guillaumin, Bull. Mus. Natl. Hist. Nat., sér. 2, 32: 188. 1960.
Calanthe subgen. Gastrorchis (Schltr.) Z. H. Feng, comb. & stat. nov. — Basionym: Gastrorchis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 33: 167. 1924.
Calanthe subgen. Hecabe (Raf.) Z. H. Feng, comb. & stat. nov. — Basionym: Hecabe Raf., Fl. Tellur. 4: 44. 1838. — Synonym: Paraphaius J. W. Zhai & F. W. Xing, Molec. Phylogen. Evol. 77: 221. 2014.
Calanthe subgen. Phaius (Lour.) Z. H. Feng, comb. & stat. nov. — Basionym: Phaius Lour., Fl. Cochinch. 2: 517, 529. 1790.
Calanthe subgen. Preptanthe (Rchb. f.) Schltr. (1912).
Calanthe subgen. Styloglossum Kakadzu (1994).
Rosaceae
Docynia Decne.
Remark: The Docynia-Docyniopsis clade has a conflicting phylogenetic position in the nuclear and plastid trees, indicating that the evolutionary relationships are not clear-cut. Liu et al. argued that “narrow generic concepts may be impractical for use”, however, this does not necessarily justify lumping because taxonomic circumscription should consider the broader implications for conservation, ecology, and evolution, simply lumping Docynia into Malus based on obviously unconvincing evidence may benifit the nomenclature but disrupt the understanding of the unique evolutionary adaptations and ecological roles of Docynia. Docynia is here kept as an independent genus, and is divided into two sections based on morpho-phylogenetic evidences: sect. Docynia encompasses D. delavayi, D. indica and D. longiunguis, and sect. Paradocynia contains only D. doumeri.
Docynia sect. Docynia
Docynia sect. Paradocynia Su Liu ex Z. H. Feng, sect. nov. Type: Docynia doumeri (Bois) C. K. Schneid.
Diagnosis: Lamina oblong to ovate, base rounded or cuneate, apex acute or acuminate, pubescent when young then glabrescent, or glabrous; styles 5, pubescent at base; pome ca 2.5‒5.5 cm in diameter, subglobose, yellow to red. Monotypic.
Eriolobus (DC.) M. Roem.
Remark: Although Liu et al. argued that the transferring of Docynia to the genus Eriolobus is untenable, they, unfortunately, did not take the similar choice as another line of evidence to keep Eriolobus apart from Malus s. s. According to Wang et al. and their previous research, Malus is dyphyletic, the North American crab apples clade being distant to the type clade, proving that it is an irrational choice to oversimply lumping Eriolobus into Malus. Eriolobus is here recognized as independent genus and is divided into three sections based on morpho-phylogenetic evidences, sect. Eriolobus includes only the type species E. trilobatus, sect. Chloromeles contains E. angustifolius, E. coronarius and E. ioensis, while sect. Florentinae solely embraces E. florentinus.
Eriolobus sect. Chloromeles (Decne.) Su Liu ex Z. H. Feng, comb. & stat. nov. — Basionym: Malus subgen. Chloromeles Decne., Nouv. Arch. Mus. Hist. Nat. Paris 10: 155. 1874. Type: Eriolobus angustifolius (Aiton) Su Liu ex Z. H. Feng.
Eriolobus sect. Eriolobus
Eriolobus sect. Florentinae (Rehder) Su Liu ex Z. H. Feng, comb. & stat. nov. — Basionym: Malus subsect. Florentinae Rehder, J. Arnold Arbor. 2: 48. 1920. Type: Eriolobus florentinus (Zuccagni) Stapf.
3.2. New Nothoranks of Pteridophytes and Spermatophytes
Amaranthaceae
Amaranthus L.
Amaranthus nothosect. Blorion Z. H. Feng, nothosect. nov. = Amaranthus sect. Blitopsis Dumort. × Amaranthus sect. Pentamorion (Beck) Mosyakin & K. R. Robertson.
Amaranthus nothosect. Pentidium Z. H. Feng, nothosect. nov. = Amaranthus sect. Pentamorion (Beck) Mosyakin & K. R. Robertson × Amaranthus sect. Pyxidium Moq.
Amaranthus nothosect. Pyxopsis Z. H. Feng, nothosect. nov. = Amaranthus sect. Blitopsis Dumort. × Amaranthus sect. Pyxidium Moq.
Amaranthus nothosubgen. Amarida Z. H. Feng, nothosubgen. nov. = Amaranthus subgen. Amaranthus × Amaranthus subgen. Acnida (L.) Aellen ex K. R. Robertson.
Amaranthus nothosubsect. Hybramaranthus Z. H. Feng, nothosubsect. nov. = Amaranthus subsect. Amaranthus × Amaranthus subsect. Hybrida Mosyakin & K. R. Robertson.
Amaranthus nothosubsect. Hynida Z. H. Feng, nothosubsect. nov. = Amaranthus sect. Acnida (L.) Mosyakin & K. R. Robertson × Amaranthus subsect. Hybrida Mosyakin & K. R. Robertson.
Amaryllidaceae
Galanthus L.
Galanthus nothoser. Woropini Z. H. Feng, nothoser. nov. = Galanthus ser. Alpini Z. H. Feng × Galanthus ser. Woronowii Kem.-Nath.
Anemiaceae
Anemia Sw.
Anemia nothosect. Anemutae Z. H. Feng, nothosect. nov. = Anemia sect. Anemia × Anemia sect. Hirsutae Prantl.
Anemia nothosect. Aninae Z. H. Feng, nothosect. nov. = Anemia sect. Anemia × Anemia sect. Collinae Prantl.
Anemia nothosect. Cosutae Z. H. Feng, nothosect. nov. = Anemia sect. Collinae Prantl. × Anemia sect. Hirsutae Prantl.
Anemia nothosect. Tomemia Z. H. Feng, nothosect. nov. = Anemia sect. Anemia × Anemia sect. Tomentosae Prantl.
Asparagaceae
Agave L.
Agave nothosect. Deslorae Z. H. Feng, nothosect. nov. = Agave sect. Deserticolae (Trel.) R. H. Webb & G. D. Starr × Agave sect. Parviflorae Hochstätter.
Agave nothosect. Hetemoratae Z. H. Feng, nothosect. nov. = Agave sect. Heteracanthae Salm-Dyck × Agave sect. Marmoratae (A. Berger) Thiede & Gideon F. Sm.
Agave nothosect. Heticracanthae Z. H. Feng, nothosect. nov. = Agave sect. Heteracanthae Salm-Dyck × Agave sect. Micracanthae Salm-Dyck.
Agave nothosect. Hibacanthae Z. H. Feng, nothosect. nov. = Agave sect. Heteracanthae Salm-Dyck × Agave sect. Hibernicae Hochstätter.
Agave nothosect. Littanthae Z. H. Feng, nothosect. nov. = Agave sect. Littaea (Tagl.) Benth. × Agave sect. Micracanthae Salm-Dyck.
Agave nothosect. Micragave Z. H. Feng, nothosect. nov. = Agave sect. Agave × Agave sect. Micracanthae Salm-Dyck.
Agave nothosect. Parranthae Z. H. Feng, nothosect. nov. = Agave sect. Heteracanthae Salm-Dyck × Agave sect. Parryanae Hochstätter.
Agave nothosect. Parvipalae Z. H. Feng, nothosect. nov. = Agave sect. Ditepalae Hochstätter × Agave sect. Parviflorae Hochstätter.
Agave nothoser. Bravanthes (Cif. & Giacom.) Z. H. Feng, comb. & stat. nov. = Agave ser. Bravoa (Lex.) Thiede & Gideon F. Sm. × Agave ser. Polianthes (L.) Thiede & Gideon F. Sm. — Basionym: × Bravanthes Cif. & Giacom., Nomencl. Fl. Ital. Pt. 1: 134. 1950.
Agave nothosubgen. Agavaea Z. H. Feng, nothosubgen. nov. = Agave subgen. Agave × Agave subgen. Littaea (Tagl.) Baker.
Asphodelaceae
Hemerocallis L.
Hemerocallis nothosect. Hemerulvae Z. H. Feng & H. N. Sun, nothosect. nov. = Hemerocallis sect. Fulvae Nakai × Hemerocallis sect. Hemerocallis.
Asteraceae
Arctium L.
Arctium nothosect. Pecarctium Z. H. Feng, nothosect. nov. = Arctium sect. Pectinata (C. Winkl.) S. López, Romasch., Susanna & N. Garcia × Arctium sect. Pseudarctium (Juz.) Duist.
Artemisia L.
Artemisia nothosubgen. Artinthium Z. H. Feng, nothosubgen. nov. = Artemisia subgen. Absinthium (Mill.) Less. × Artemisia subgen. Artemisia.
Artemisia nothosubgen. Dracunthium Z. H. Feng, nothosubgen. nov. = Artemisia subgen. Absinthium (Mill.) Less. × Artemisia subgen. Dracunculus (Besser) Rydb.
Artemisia nothosubgen. Ponculus Z. H. Feng, nothosubgen. nov. = Artemisia subgen. Dracunculus (Besser) Rydb. × Artemisia subgen. Ponticae B. H. Jiao ex T. G. Gao.
Artemisia nothosubgen. Sesinthium Z. H. Feng, nothosubgen. nov. = Artemisia subgen. Absinthium (Mill.) Less. × Artemisia subgen. Seriphidium Besser ex Less.
Baccharis L.
Baccharis nothosect. Pedifoliae Z. H. Feng, nothosect. nov. = Baccharis sect. Cuneifoliae DC. × Baccharis sect. Pedicellatae Heering.
Cousinia Cass.
Cousinia nothosect. Alpiopsis Z. H. Feng, nothosect. nov. = Cousinia sect. Alpinae Bunge × Cousinia sect. Jurineopsis (Juz.) Tscherneva.
Cousinia nothosect. Coronopsis Z. H. Feng, nothosect. nov. = Cousinia sect. Coronophora (Juz.) Rech. f. × Cousinia sect. Jurineopsis (Juz.) Tscherneva.
Cousinia nothosect. Cynarocaules Z. H. Feng, nothosect. nov. = Cousinia sect. Cynaroideae Bunge × Cousinia sect. Leiocaules Bunge.
Cousinia nothosect. Cynaroferae Z. H. Feng, nothosect. nov. = Cousinia sect. Cynaroideae Bunge × Cousinia sect. Pugioniferae Bunge.
Cousinia nothosect. Pulchecarpae Z. H. Feng, nothosect. nov. = Cousinia sect. Microcarpae Bunge × Cousinia sect. Pulchellae Rech. f.
Cousinia nothosect. Pulchepinae Z. H. Feng, nothosect. nov. = Cousinia sect. Alpinae Bunge × Cousinia sect. Pulchellae Rech. f.
Cousinia nothosect. Serracousinia Z. H. Feng, nothosect. nov. = Cousinia sect. Eriocousinia Tscherneva × Cousinia sect. Serratuloides Bunge.
Cousinia nothosect. Sphaeroides Z. H. Feng, nothosect. nov. = Cousinia sect. Serratuloides Bunge × Cousinia sect. Sphaerocephalae Bunge.
Liatris Gaertn. ex Schreb.
Liatris nothosect. Gramago Z. H. Feng, nothosect. nov. = Liatris sect. Graminifolium G. L. Nesom × Liatris sect. Suprago (Gaertn.) DC.
Liatris nothosect. Liafolium Z. H. Feng, nothosect. nov. = Liatris sect. Graminifolium G. L. Nesom × Liatris sect. Liatris.
Liatris nothosect. Liprago Z. H. Feng, nothosect. nov. = Liatris sect. Liatris × Liatris sect. Suprago G. L. Nesom.
Liatris nothosect. Lirago Z. H. Feng, nothosect. nov. = Liatris sect. Liatris × Liatris sect. Vorago G. L. Nesom.
Liatris nothoser. Elerago Z. H. Feng, nothoser. nov. = Liatris ser. Elegantes (Alexander) Gaiser ex G. L. Nesom × Liatris sect. Vorago G. L. Nesom.
Liatris nothoser. Liprago (Z. H. Feng) Z. H. Feng, stat. nov. = Liatris ser. Liatris × Liatris sect. Suprago G. L. Nesom. — Basionym: Liatris nothosect. Liprago Z. H. Feng, this publication.
Liatris nothoser. Lirago (Z. H. Feng) Z. H. Feng, stat. nov. = Liatris ser. Liatris × Liatris sect. Vorago G. L. Nesom. — Basionym: Liatris nothosect. Lirago Z. H. Feng, this publication.
Liatris nothoser. Punosae Z. H. Feng, nothoser. nov. = Liatris ser. Punctatae (Alexander) Gaiser ex Fernald × Liatris ser. Scariosae (Alexander) Gaiser ex Fernald.
Liatris nothoser. Scrago Z. H. Feng, nothoser. nov. = Liatris ser. Scariosae (Alexander) Gaiser ex Fernald × Liatris sect. Suprago (Gaertn.) DC.
Athyriaceae
Athyrium Roth
Athyrium nothosect. Bistichoides Z. H. Feng, nothosect. nov. = Athyrium sect. Biserrulata (Ching & Y. T. Hsieh) R. Wei & X. C. Zhang × Athyrium sect. Polystichoides Ching & Y. T. Hsieh.
Athyrium nothosect. Otoniana Z. H. Feng, nothosect. nov. = Athyrium sect. Mackinnoniana (Ching & Y. T. Hsieh) Z. R. Wang × Athyrium sect. Otophora Z. R. Wang.
Athyrium nothosect. Otoscentia Z. H. Feng, nothosect. nov. = Athyrium sect. Otophora Z. R. Wang × Athyrium sect. Yokoscentia (Z. R. Wang) R. Wei & X. C. Zhang.
Athyrium nothosect. Polyphora Z. H. Feng, nothosect. nov. = Athyrium sect. Otophora Z. R. Wang × Athyrium sect. Polystichoides Ching & Y. T. Hsieh.
Athyrium nothosect. Yokoniana Z. H. Feng, nothosect. nov. = Athyrium sect. Mackinnoniana (Ching & Y. T. Hsieh) Z. R. Wang × Athyrium sect. Yokoscentia (Z. R. Wang) R. Wei & X. C. Zhang.
Deparia Medik.
Deparia nothosect. Lunoathyrium Z. H. Feng, nothosect. nov. = Deparia sect. Dryoathyrium (Ching) M. Kato × Deparia sect. Lunathyrium (Koidz.) M. Kato.
Deparia nothosubsect. Depopsis Z. H. Feng, nothosubsect. nov. = Deparia subsect. Athyriopsis (Ching) L. Y. Kuo, M. Kato & W. L. Chiou × Deparia subsect. Deparia.
Deparia nothosubsect. Lunathyriopsis Z. H. Feng, nothosubsect. nov. = Deparia subsect. Athyriopsis (Ching) L. Y. Kuo, M. Kato & W. L. Chiou × Deparia sect. Lunathyrium (Koidz.) M. Kato.
Diplazium Sw.
Diplazium nothosect. Anisostegia Z. H. Feng, nothosect. nov. = Diplazium sect. Anisogonium (C. Presl) Alderw. × Diplazium sect. Dolichostegia (W. M. Chu & Z. R. He) R. Wei.
Diplazium nothosect. Anisotodia Z. H. Feng, nothosect. nov. = Diplazium sect. Allantodia (R. Br.) R. Wei × Diplazium sect. Anisogonium (C. Presl) Alderw.
Diplazium nothosect. Dolichomatum Z. H. Feng, nothosect. nov. = Diplazium sect. Dolichostegia (W. M. Chu & Z. R. He) R. Wei × Diplazium sect. Longirhizomatum (W. M. Chu & Z. R. He) R. Wei.
Diplazium nothosect. Sibistegia Z. H. Feng, nothosect. nov. = Diplazium subgen. Sibirica R. Wei & X. C. Zhang × Diplazium sect. Dolichostegia (W. M. Chu & Z. R. He) R. Wei.
Diplazium nothosubgen. Sibipteris Z. H. Feng, nothosubgen. nov. = Diplazium subgen. Callipteris (Bory) R. Wei & X. C. Zhang × Diplazium subgen. Sibirica R. Wei & X. C. Zhang.
Berberidaceae
Epimedium L.
Epimedium nothosect. Rhizepimedium (Z. H. Feng) Z. H. Feng, stat. nov. = Epimedium sect. Epimedium × Epimedium subgen. Rhizophyllum (Fisch. & C. A. Mey.) Stearn. — Basionym: Epimedium nothosubgen. Rhizepimedium Z. H. Feng, this publication.
Epimedium nothosect. Rhizoceras Z. H. Feng, nothosect. nov. = Epimedium sect. Macroceras C. Morren & Decne. × Epimedium subgen. Rhizophyllum (Fisch. & C. A. Mey.) Stearn.
Epimedium nothosect. Macrepimedium Z. H. Feng, nothosect. nov. = Epimedium sect. Epimedium × Epimedium sect. Macroceras C. Morren & Decne.
Epimedium nothoser. Dolidianae Z. H. Feng, nothoser. nov. = Epimedium ser. Dolichocera Stearn × Epimedium ser. Davidianae Stearn.
Epimedium nothosubgen. Rhizepimedium Z. H. Feng, nothosubgen. nov. = Epimedium subgen. Epimedium × Epimedium subgen. Rhizophyllum (Fisch. & C. A. Mey.) Stearn.
Cistaceae
Helianthemum Mill.
Helianthemum nothosect. Lavanthemum Z. H. Feng, nothosect. nov. = Helianthemum sect. Helianthemum × Helianthemum sect. Lavandulaceum G. López.
Helianthemum nothosect. Pseudanthemum Z. H. Feng, nothosect. nov. = Helianthemum sect. Helianthemum × Helianthemum sect. Pseudocistus Dunal.
Helianthemum nothosubgen. Erionthemum Z. H. Feng, nothosubgen. nov. = Helianthemum subgen. Eriocarpum (Dunal) Martín-Hernanz, Velayos, Albaladejo & Aparicio × Helianthemum subgen. Helianthemum.
Helianthemum nothosubgen. Plecanthemum Z. H. Feng, nothosubgen. nov. = Helianthemum subgen. Helianthemum × Helianthemum subgen. Plectolobum Willk.
Crassulaceae
Aichryson Webb & Berthel.
Aichryson nothosect. Aichrobia Z. H. Feng, nothosect. nov. = Aichryson sect. Aichryson × Aichryson sect. Macrobia Webb & Berthel.
Kalanchoe Adans.
Kalanchoe nothoser. Frutisores Z. H. Feng, nothoser. nov. = Kalanchoe ser. Fruticosa Gideon F. Sm. × Kalanchoe ser. Invasores Gideon F. Sm.
Kalanchoe nothoser. Herilana Z. H. Feng, nothoser. nov. = Kalanchoe ser. Herbacea Gideon F. Sm. × Kalanchoe ser. Vilana Gideon F. Sm.
Kalanchoe nothoser. Hersores Z. H. Feng, nothoser. nov. = Kalanchoe ser. Herbacea Gideon F. Sm. × Kalanchoe ser. Invasores Gideon F. Sm.
Kalanchoe nothoser. Invalana Z. H. Feng, nothoser. nov. = Kalanchoe ser. Invasores Gideon F. Sm. × Kalanchoe ser. Vilana Gideon F. Sm.
Petrosedum Grulich
Remark: Afferni assigned the genotype Petrosedum rupestre to Petrosedum ser. Reflexa, according to Art. 22.1 of ICN , the type serie must be named as Petrosedum ser. Petrosedum.
Petrosedum nothoser. Petrorecta Z. H. Feng, nothoser. nov. = Petrosedum ser. Petrosedum × Petrosedum ser. Recta Afferni.
Petrosedum nothoser. Petudorecta Z. H. Feng, nothoser. nov. = Petrosedum ser. Petrosedum × Petrosedum ser. Pseudorecta Afferni.
Petrosedum nothoser. Reseudorecta Z. H. Feng, nothoser. nov. = Petrosedum ser. Pseudorecta Afferni × Petrosedum ser. Recta Afferni.
Dryopteridaceae
Dryopteris Adans.
Dryopteris nothosect. Aedium Z. H. Feng, nothosect. nov. = Dryopteris sect. Aemulae Fraser-Jenk. × Dryopteris subgen. Lophodium (Newman) Tzvelev.
Dryopteris nothosect. Aemotae Z. H. Feng, nothosect. nov. = Dryopteris sect. Aemulae Fraser-Jenk. × Dryopteris sect. Remotae Fraser-Jenk.
Dryopteris nothosect. Aepteris Z. H. Feng, nothosect. nov. = Dryopteris sect. Aemulae Fraser-Jenk. × Dryopteris sect. Dryopteris.
Dryopteris nothosect. Aevariae Z. H. Feng, nothosect. nov. = Dryopteris sect. Aemulae Fraser-Jenk. × Dryopteris sect. Erythrovariae H. Ito.
Dryopteris nothosect. Drillodium Z. H. Feng, nothosect. nov. = Dryopteris sect. Dryopteris × Dryopteris sect. Fibrillosae Ching × Dryopteris subgen. Lophodium (Newman) Tzvelev.
Dryopteris nothosect. Dryllidae Z. H. Feng, nothosect. nov. = Dryopteris sect. Dryopteris × Dryopteris sect. Pallidae Fraser-Jenk.
Dryopteris nothosect. Dryollosae Z. H. Feng, nothosect. nov. = Dryopteris sect. Dryopteris × Dryopteris sect. Fibrillosae Ching.
Dryopteris nothosect. Dryphodium (Z. H. Feng) Z. H. Feng, stat. nov. = Dryopteris sect. Dryopteris × Dryopteris subgen. Lophodium (Newman) Tzvelev. — Basionym: Dryopteris nothosubgen. Dryphodium Z. H. Feng, this publication.
Dryopteris nothosect. Erypallidae Z. H. Feng, nothosect. nov. = Dryopteris sect. Erythrovariae H. Ito × Dryopteris sect. Pallidae Fraser-Jenk.
Dryopteris nothosect. Erypandae Z. H. Feng, nothosect. nov. = Dryopteris sect. Erythrovariae H. Ito × Dryopteris sect. Pandae Fraser-Jenk.
Dryopteris nothosect. Fidae Z. H. Feng, nothosect. nov. = Dryopteris sect. Fibrillosae Ching × Dryopteris sect. Pallidae Fraser-Jenk.
Dryopteris nothosect. Fiphodium Z. H. Feng, nothosect. nov. = Dryopteris sect. Fibrillosae Ching × Dryopteris subgen. Lophodium (Newman) Tzvelev.
Dryopteris nothosect. Hiropteris Z. H. Feng, nothosect. nov. = Dryopteris sect. Hirtipedes Fraser-Jenk. × Dryopteris subgen. Pycnopteris (Moore) Ching.
Dryopteris nothosect. Hirtidae Z. H. Feng, nothosect. nov. = Dryopteris sect. Hirtipedes Fraser-Jenk. × Dryopteris sect. Pallidae Fraser-Jenk.
Dryopteris nothosect. Hirtodon Z. H. Feng, nothosect. nov. = Dryopteris sect. Diclisodon (T. Moore) C. Chr. × Dryopteris sect. Hirtipedes Fraser-Jenk.
Dryopteris nothosect. Hirtosae Z. H. Feng, nothosect. nov. = Dryopteris sect. Fibrillosae Ching × Dryopteris sect. Hirtipedes Fraser-Jenk.
Dryopteris nothosect. Lolidae Z. H. Feng, nothosect. nov. = Dryopteris subgen. Lophodium (Newman) Tzvelev × Dryopteris sect. Pallidae Fraser-Jenk.
Dryopteris nothosect. Lophatae Z. H. Feng, nothosect. nov. = Dryopteris subgen. Lophodium (Newman) Tzvelev × Dryopteris sect. Marginatae Fraser-Jenk.
Dryopteris nothosect. Margisodon Z. H. Feng, nothosect. nov. = Dryopteris sect. Diclisodon (T. Moore) C. Chr. × Dryopteris sect. Marginatae Fraser-Jenk.
Dryopteris nothosect. Pallindae Z. H. Feng, nothosect. nov. = Dryopteris sect. Pallidae Fraser-Jenk. × Dryopteris sect. Pandae Fraser-Jenk.
Dryopteris nothosect. Palopteris Z. H. Feng, nothosect. nov. = Dryopteris sect. Pallidae Fraser-Jenk. × Dryopteris subgen. Pycnopteris (Moore) Ching.
Dryopteris nothosect. Panphodium Z. H. Feng, nothosect. nov. = Dryopteris subgen. Lophodium (Newman) Tzvelev × Dryopteris sect. Pandae Fraser-Jenk.
Dryopteris nothosect. Panpteris Z. H. Feng, nothosect. nov. = Dryopteris sect. Dryopteris × Dryopteris sect. Pandae Fraser-Jenk.
Dryopteris nothosect. Remolosae Z. H. Feng, nothosect. nov. = Dryopteris sect. Fibrillosae Ching × Dryopteris sect. Remotae Fraser-Jenk.
Dryopteris nothosect. Remophodium Z. H. Feng, nothosect. nov. = Dryopteris subgen. Lophodium (Newman) Tzvelev × Dryopteris sect. Remotae Fraser-Jenk.
Dryopteris nothosect. Remopteris Z. H. Feng, nothosect. nov. = Dryopteris sect. Dryopteris × Dryopteris sect. Remotae Fraser-Jenk.
Dryopteris nothosect. Shiopsis Z. H. Feng, nothosect. nov. = Dryopteris sect. Dryopsis (Holttum & P. J. Edwards) Li Bing Zhang × Dryopteris sect. Shiehia L. Y. Kuo & Y. H. Chang.
Dryopteris nothosect. Vavariae Z. H. Feng, nothosect. nov. = Dryopteris sect. Erythrovariae H. Ito × Dryopteris sect. Variae Fraser-Jenk.
Dryopteris nothosubgen. Dryphodium Z. H. Feng, nothosubgen. nov. = Dryopteris subgen. Dryopteris × Dryopteris subgen. Lophodium (Newman) Tzvelev.
Dryopteris nothosubgen. Erythteris Z. H. Feng, nothosubgen. nov. = Dryopteris subgen. Dryopteris × Dryopteris subgen. Erythrovariae (H. Ito) Fraser-Jenk.
Dryopteris nothosubgen. Lophema Z. H. Feng, nothosubgen. nov. = Dryopteris subgen. Lophodium (Newman) Tzvelev × Dryopteris subgen. Nothoperanema Tagawa.
Dryopteris nothosubgen. Notheris Z. H. Feng, nothosubgen. nov. = Dryopteris subgen. Dryopteris × Dryopteris subgen. Nothoperanema Tagawa.
Dryopteris nothosubgen. Nothovariae Z. H. Feng, nothosubgen. nov. = Dryopteris subgen. Erythrovariae (H. Ito) Fraser-Jenk. × Dryopteris subgen. Nothoperanema Tagawa.
Dryopteris nothosubgen. Pycyopteris Z. H. Feng, nothosubgen. nov. = Dryopteris subgen. Dryopteris × Dryopteris subgen. Pycnopteris (Moore) Ching.
Heliconiaceae
Heliconia L.
Heliconia nothosect. Epiconia Z. H. Feng, nothosect. nov. = Heliconia sect. Episcopales L. Anderss. × Heliconia sect. Heliconia.
Heliconia nothosect. Epistrobus Z. H. Feng, nothosect. nov. = Heliconia sect. Episcopales L. Anderss. × Heliconia sect. Taeniostrobus Kuntze.
Heliconia nothosubgen. Helitrobus Z. H. Feng, nothosubgen. nov. = Heliconia subgen. Heliconia × Heliconia subgen. Taeniostrobus (Kuntze) Griggs.
Hydrangeaceae
Hydrangea Gronov.
Hydrangea nothosect. Dichophyllae Z. H. Feng, nothosect. nov. = Hydrangea sect. Dichroa (Lour.) Y. De Smet & Samain × Hydrangea sect. Macrophyllae (E. M. McClint.) Y. De Smet & Samain.
Hydrangea nothosect. Hirtophyllae Z. H. Feng, nothosect. nov. = Hydrangea sect. Hirtae Y. De Smet & Samain × Hydrangea sect. Macrophyllae (E. M. McClint.) Y. De Smet & Samain.
Hydrangea nothosect. Kawirtae Z. H. Feng, nothosect. nov. = Hydrangea sect. Hirtae Y. De Smet & Samain × Hydrangea sect. Kawagoeanae X. D. Yang & S. X. Yu.
Hypericaceae
Hypericum L.
Hypericum nothosect. Drosoricum Z. H. Feng, nothosect. nov. = Hypericum sect. Drosocarpium Spach × Hypericum sect. Hypericum.
Hypericum nothoser. Drosoricum (Z. H. Feng) Z. H. Feng, stat. nov. = Hypericum sect. Drosocarpium Spach × Hypericum ser. Hypericum. — Basionym: Hypericum nothosect. Drosoricum Z. H. Feng, this publication.
Hypericum nothosubsect. Aethiosepalum Z. H. Feng, nothosubsect. nov. = Hypericum subsect. Adenosepalum N. Robson × Hypericum subsect. Aethiopica N. Robson.
Hypericum nothosubsect. Drosoricum (Z. H. Feng) Z. H. Feng, stat. nov. = Hypericum sect. Drosocarpium Spach × Hypericum subsect. Hypericum. — Basionym: Hypericum nothosect. Drosoricum Z. H. Feng, this publication.
Iridaceae
Gladiolus L.
Gladiolus nothosect. Homifolii Z. H. Feng, nothosect. nov. = Gladiolus sect. Homoglossum (Salisb.) Goldblatt & J. C. Manning × Gladiolus sect. Linearifolii (M. P. de Vos) Goldblatt & J. C. Manning.
Gladiolus nothoser. Trearifolii Z. H. Feng, nothoser. nov. = Gladiolus ser. Linearifolii Goldblatt & J. C. Manning × Gladiolus ser. Tristes Goldblatt & J. C. Manning.
Lamiaceae
Lavandula L.
Lavandula nothosect. Laventatae Z. H. Feng, nothosect. nov. = Lavandula sect. Dentatae Suárez-Cerv. & Seoane-Camba × Lavandula sect. Lavandula.
Origanum L.
Origanum nothosect. Amilamentum Z. H. Feng, nothosect. nov. = Origanum sect. Amaracus (Gled.) Vogel × Origanum sect. Brevifilamentum Ietsw.
Origanum nothosect. Amolicon Z. H. Feng, nothosect. nov. = Origanum sect. Amaracus (Gled.) Vogel × Origanum sect. Anatolicon Ietsw.
Origanum nothosect. Anatorana Z. H. Feng, nothosect. nov. = Origanum sect. Anatolicon Ietsw. × Origanum sect. Majorana (Mill.) Benth.
Origanum nothosect. Anatoriganum Z. H. Feng, nothosect. nov. = Origanum sect. Anatolicon Ietsw. × Origanum sect. Origanum.
Origanum nothosect. Brevicorolla Z. H. Feng, nothosect. nov. = Origanum sect. Brevifilamentum Ietsw. × Origanum sect. Prolaticorolla Ietsw.
Origanum nothosect. Brevifilicon Z. H. Feng, nothosect. nov. = Origanum sect. Anatolicon Ietsw. × Origanum sect. Brevifilamentum Ietsw.
Origanum nothosect. Eloticorolla Z. H. Feng, nothosect. nov. = Origanum sect. Elongatispica Ietsw. × Origanum sect. Prolaticorolla Ietsw.
Origanum nothosect. Longicorolla Z. H. Feng, nothosect. nov. = Origanum sect. Longitubus Ietsw. × Origanum sect. Prolaticorolla Ietsw.
Origanum nothosect. Origalyx Z. H. Feng, nothosect. nov. = Origanum sect. Chilocalyx (Briq.) Ietsw. × Origanum sect. Origanum.
Origanum nothosect. Origanomajorana (Domin) Z. H. Feng, comb. & stat. nov. = Origanum sect. Majorana (Mill.) Benth. × Origanum sect. Origanum. — Basionym: × Origanomajorana Domin, Preslia 13-15: 197. 1935.
Origanum nothosect. Orimentum Z. H. Feng, nothosect. nov. = Origanum sect. Brevifilamentum Ietsw. × Origanum sect. Origanum.
Origanum nothosect. Oriracus Z. H. Feng, nothosect. nov. = Origanum sect. Amaracus (Gled.) Vogel × Origanum sect. Origanum.
Origanum nothosect. Protolicon Z. H. Feng, nothosect. nov. = Origanum sect. Anatolicon Ietsw. × Origanum sect. Prolaticorolla Ietsw.
Teucrium L.
Teucrium nothosect. Chamaepolium Z. H. Feng, nothosect. nov. = Teucrium sect. Chamaedrys (Mill.) Schreb. × Teucrium sect. Polium (Mill.) Schreb.
Teucrium nothosect. Monpolium Z. H. Feng, nothosect. nov. = Teucrium sect. Montana Lázaro Ibiza × Teucrium sect. Polium (Mill.) Schreb.
Teucrium nothosect. Stachyodonia Z. H. Feng, nothosect. nov. = Teucrium sect. Scorodonia (Hill) Schreb. × Teucrium sect. Stachyobotrys Benth.
Teucrium nothosubsect. Canchamaedrys Z. H. Feng, nothosubsect. nov. = Teucrium subsect. Cana Melnikov × Teucrium subsect. Chamaedrys (Mill.) Kästner.
Teucrium nothosubsect. Chamaepolium (Z. H. Feng) Z. H. Feng, stat. nov. = Teucrium subsect. Chamaedrys (Mill.) Kästner × Teucrium subsect. Polium (Mill.) Kästner. — Basionym: Teucrium nothosect. Chamaepolium Z. H. Feng, this publication.
Teucrium nothosubsect. Monfolia Z. H. Feng, nothosubsect. nov. = Teucrium sect. Montana Lázaro Ibiza × Teucrium subsect. Rotundifolia M. E. Cohen.
Teucrium nothosubsect. Montolium Z. H. Feng, nothosubsect. nov. = Teucrium sect. Montana Lázaro Ibiza × Teucrium subsect. Polium (Mill.) Kästner.
Teucrium nothosubsect. Polifolia Z. H. Feng, nothosubsect. nov. = Teucrium subsect. Polium (Mill.) Kästner × Teucrium subsect. Rotundifolia M. E. Cohen.
Teucrium nothosubsect. Pomilum Z. H. Feng, nothosubsect. nov. = Teucrium subsect. Polium (Mill.) Kästner × Teucrium subsect. Pumilum Rivas Mart.
Teucrium nothosubsect. Popilosa Z. H. Feng, nothosubsect. nov. = Teucrium subsect. Polium (Mill.) Kästner × Teucrium subsect. Simplicipilosa S. Puech.
Teucrium nothosubsect. Pupilosa Z. H. Feng, nothosubsect. nov. = Teucrium subsect. Pumilum Rivas Mart. × Teucrium subsect. Simplicipilosa S. Puech.
Teucrium nothosubsect. Rotundipilosa Z. H. Feng, nothosubsect. nov. = Teucrium subsect. Rotundifolia M. E. Cohen × Teucrium subsect. Simplicipilosa S. Puech.
Lentibulariaceae
Pinguicula L.
Pinguicula nothosect. Micula Z. H. Feng, nothosect. nov. = Pinguicula sect. Micranthus Casper × Pinguicula sect. Pinguicula.
Pinguicula nothosubgen. Pinoceras Z. H. Feng, nothosubgen. nov. = Pinguicula subgen. Pinguicula × Pinguicula subgen. Temnoceras Barnhart.
Marattiaceae
Danaea Sm.
Danaea nothosubgen. Dalodanaea Z. H. Feng, nothosubgen. nov. = Danaea subgen. Danaea × Danaea subgen. Holodanaea C. Presl.
Danaea nothosubgen. Darodanaea Z. H. Feng, nothosubgen. nov. = Danaea subgen. Arthrodanaea C. Presl × Danaea subgen. Danaea.
Musaceae
Musa L.
Musa nothosect. Mulimusa Z. H. Feng, nothosect. nov. = Musa sect. Australimusa Cheesman × Musa sect. Musa.
Nymphaeaceae
Nymphaea L.
Nymphaea nothosect. Nymtha Zi Jun Li & Z. H. Feng, nothosect. nov. = Nymphaea sect. Nymphaea × Nymphaea sect. Xanthantha (Casp.) Wiersema.
Nymphaea nothosubgen. Aneceras Zi Jun Li & Z. H. Feng, nothosubgen. nov. = Nymphaea subgen. Anecphya (Casp.) Conard × Nymphaea subgen. Brachyceras (Casp.) Conard. (Figure 1)
Nymphaea nothosubgen. Anelotos Zi Jun Li & Z. H. Feng, nothosubgen. nov. = Nymphaea subgen. Anecphya (Casp.) Conard × Nymphaea subgen. Lotos (DC.) Conard. (Figure 1)
Nymphaea nothosubgen. Brachynymphaea Zi Jun Li & Z. H. Feng, nothosubgen. nov. = Nymphaea subgen. Brachyceras (Casp.) Conard × Nymphaea subgen. Nymphaea. (Figure 1)
Orchidaceae
Anacamptis Rich.
Anacamptis nothosect. Anacranthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Anacamptis × Anacamptis sect. Andranthus (Schltr.) F. M. Vázquez. — Synonym: × Anacampaludorchis P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Anacamptis nothosect. Andorianthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Andranthus (Schltr.) F. M. Vázquez × Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov.
Anacamptis nothosect. Andraenanthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Andranthus (Schltr.) F. M. Vázquez × Anacamptis sect. Phalaenanthus (Schltr.) Kuropatkin & Efimov.
Anacamptis nothosect. Andriopanthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Andranthus (Schltr.) F. M. Vázquez × Anacamptis sect. Coriophorae (Parl.) P. Quentin × Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov.
Anacamptis nothosect. Andriophorae Z. H. Feng, nothosect. nov. = Anacamptis sect. Andranthus (Schltr.) F. M. Vázquez × Anacamptis sect. Coriophorae (Parl.) P. Quentin. — Synonym: × Anteriopaludorchis P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Anacamptis nothosect. Andryae Z. H. Feng, nothosect. nov. = Anacamptis sect. Andranthus (Schltr.) F. M. Vázquez × Anacamptis sect. Boryae H. Kretzschmar, Eccarius & H. Dietr.
Anacamptis nothosect. Anlaenanthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Anacamptis × Anacamptis sect. Phalaenanthus (Schltr.) Kuropatkin & Efimov. — Synonym: × Anacampteulenia P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Anacamptis nothosect. Anorianthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Anacamptis × Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov. — Synonym: × Anacampterorchis P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Anacamptis nothosect. Anoriophorae Z. H. Feng, nothosect. nov. = Anacamptis sect. Anacamptis × Anacamptis sect. Coriophorae (Parl.) P. Quentin. — Synonym: × Anteriocamptis P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Figure 1. Representative cultivars from three Nymphaea nothosubgenera: A-B. Nymphaea ‘New Poem of the Nile’ (Nymphaea nothosubgen. Aneceras), C-D. Nymphaea ‘Tiwa Watree’ (Nymphaea nothosubgen. Anelotos), E-F. Nymphaea ‘Detective Erika’, G-H. Nymphaea ‘Siam Pink 1’ (Nymphaea nothosubgen. Brachynymphaea). All photos by Zi Jun Li.
Anacamptis nothosect. Boraenanthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Boryae H. Kretzschmar, Eccarius & H. Dietr. × Anacamptis sect. Phalaenanthus (Schltr.) Kuropatkin & Efimov.
Anacamptis nothosect. Borianthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Boryae H. Kretzschmar, Eccarius & H. Dietr. × Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov.
Anacamptis nothosect. Borophorae Z. H. Feng, nothosect. nov. = Anacamptis sect. Boryae H. Kretzschmar, Eccarius & H. Dietr. × Anacamptis sect. Coriophorae (Parl.) P. Quentin.
Anacamptis nothosect. Colaenanthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Coriophorae (Parl.) P. Quentin × Anacamptis sect. Phalaenanthus (Schltr.) Kuropatkin & Efimov. — Synonym: × Anteriomeulenia P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Anacamptis nothosect. Corianthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Coriophorae (Parl.) P. Quentin × Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov. — Synonym: × Anterioherorchis P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Anacamptis nothosect. Moraccatae Z. H. Feng, nothosect. nov. = Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov × Anacamptis sect. Saccatae (Rchb. f.) P. Quentin.
Anacamptis nothosect. Moraenanthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov × Anacamptis sect. Phalaenanthus (Schltr.) Kuropatkin & Efimov.
Anacamptis nothosect. Phalaccatae Z. H. Feng, nothosect. nov. = Anacamptis sect. Phalaenanthus (Schltr.) Kuropatkin & Efimov × Anacamptis sect. Saccatae (Rchb. f.) P. Quentin.
Calanthe R. Br.
Calanthe nothosect. Triverna Z. H. Feng, nothosect. nov. = Calanthe sect. Tricarinata J. W. Zhai, Z. J. Liu & F. W. Xing × Calanthe sect. Verna Kakadzu.
Calanthe nothosubgen. Gastrocalanthe (A. D. Hawkes) Z. H. Feng, comb. & stat. nov. = Calanthe subgen. Calanthe × Calanthe subgen. Gastrorchis (Schltr.) Z. H. Feng. — Basionym: × Gastrocalanthe A. D. Hawkes, Orchid J. 1: 245. 1952.
Calanthe nothosubgen. Gastrophaius (A. D. Hawkes) Z. H. Feng, comb. & stat. nov. = Calanthe subgen. Gastrorchis (Schltr.) Z. H. Feng × Calanthe subgen. Phaius (Lour.) Z. H. Feng. — Basionym: × Gastrophaius A. D. Hawkes, Orchid J. 1: 245. 1952.
Calanthe nothosubgen. Phaiocalanthe (Rolfe) Z. H. Feng, comb. & stat. nov. = Calanthe subgen. Calanthe × Calanthe subgen. Phaius (Lour.) Z. H. Feng. — Basionym: × Phaiocalanthe Rolfe, J. Linn. Soc., Bot. 24: 168. 1887.
Calanthe nothosubgen. Phaiopreptanthe (Kerch.) Z. H. Feng, comb. & stat. nov. = Calanthe subgen. Phaius (Lour.) Z. H. Feng × Calanthe subgen. Preptanthe (Rchb. f.) Schltr. — Basionym: × Phaiopreptanthe Kerch., Le Liv. Orch.: 485. 1894.
Calanthe nothosubgen. Preptacalanthe (J. M. H. Shaw) Z. H. Feng, comb. & stat. nov. = Calanthe subgen. Calanthe × Calanthe subgen. Preptanthe (Rchb. f.) Schltr. — Basionym: × Preptacalanthe J. M. H. Shaw, Orchid Rev. Suppl., 121(1301): 14. 2013.
Cypripedium L.
Cypripedium nothosect. Cypetala Z. H. Feng, nothosect. nov. = Cypripedium sect. Cypripedium × Cypripedium sect. Obtusipetala (Pfitzer) P. J. Cribb.
Cypripedium nothosubsect. Cypantha Z. H. Feng, nothosubsect. nov. = Cypripedium subsect. Cypripedium × Cypripedium subsect. Macrantha (Kraenzl.) P. J. Cribb.
Cypripedium nothosubsect. Obtupedium Z. H. Feng, nothosubsect. nov. = Cypripedium subsect. Cypripedium × Cypripedium sect. Obtusipetala (Pfitzer) P. J. Cribb.
Dactylorhiza Neck. ex Nevski
Dactylorhiza nothosect. Anculatae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Angorhiza Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Dactylorhiza.
Dactylorhiza nothosect. Aristoglossum Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Aristatae Aver. × Dactylorhiza sect. Coeloglossum (Hartm.) Efimov.
Dactylorhiza nothosect. Coelofoliae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Coeloglossum (Hartm.) Efimov.
Dactylorhiza nothosect. Coelolatae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Coeloglossum (Hartm.) Efimov × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Dactycinae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Dactylorhiza × Dactylorhiza sect. Sambucinae (Parl.) Smoljan.
Dactylorhiza nothosect. Dactyloglossum (P. F. Hunt & Summerh.) Z. H. Feng, comb. & stat. nov. = Dactylorhiza sect. Coeloglossum (Hartm.) Efimov × Dactylorhiza sect. Dactylorhiza. — Basionym: × Dactyloglossum P. F. Hunt & Summerh., Watsonia 6: 132. 1965.
Dactylorhiza nothosect. Daculacinae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Dactylorhiza × Dactylorhiza sect. Maculatae (Parl.) Smoljan. × Dactylorhiza sect. Sambucinae (Parl.) Smoljan.
Dactylorhiza nothosect. Ibefoliae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Iberanthus (Schltr.) Smoljan.
Dactylorhiza nothosect. Iberatae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Iberanthus (Schltr.) Smoljan. × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Iberorhiza Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Dactylorhiza × Dactylorhiza sect. Iberanthus (Schltr.) Smoljan.
Dactylorhiza nothosect. Macoglossum Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Coeloglossum (Hartm.) Efimov × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Macucinae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Maculatae (Parl.) Smoljan. × Dactylorhiza sect. Sambucinae (Parl.) Smoljan.
Dactylorhiza nothosect. Macufoliae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Macurhiza Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Dactylorhiza × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Macustirhiza Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Dactylorhiza × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Samboglossum Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Coeloglossum (Hartm.) Efimov × Dactylorhiza sect. Sambucinae (Parl.) Smoljan.
Dactylorhiza nothosect. Samtifoliae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Sambucinae (Parl.) Smoljan.
Gymnadenia R. Br.
Gymnadenia nothosect. Balcatiana Z. H. Feng, nothosect. nov. = Gymnadenia sect. Balcanica Eccarius × Gymnadenia sect. Praegnatiana Eccarius.
Gymnadenia nothosect. Botenia Z. H. Feng, nothosect. nov. = Gymnadenia sect. Botanicana Eccarius × Gymnadenia sect. Gymnadenia.
Gymnadenia nothosect. Gygnatiana Z. H. Feng, nothosect. nov. = Gymnadenia sect. Gymnadenia × Gymnadenia sect. Praegnatiana Eccarius.
Gymnadenia nothosect. Gymneiniana Z. H. Feng, nothosect. nov. = Gymnadenia sect. Gymnadenia × Gymnadenia sect. Wettsteiniana Eccarius.
Gymnadenia nothosect. Gymnigratiana Z. H. Feng, nothosect. nov. = Gymnadenia sect. Gymnadenia × Gymnadenia sect. Nigritella (Rich.) O. Gerbaud & P. Quentin × Gymnadenia sect. Praegnatiana Eccarius.
Gymnadenia nothosect. Gymnigritella (E. G. Camus) Z. H. Feng, comb. & stat. nov. = Gymnadenia sect. Gymnadenia × Gymnadenia sect. Nigritella (Rich.) O. Gerbaud & P. Quentin. — Basionym: × Gymnigritella E. G. Camus, J. Bot. (Morot) 6: 484. 1892.
Gymnadenia nothosect. Nigratiana Z. H. Feng, nothosect. nov. = Gymnadenia sect. Nigritella (Rich.) O. Gerbaud & P. Quentin × Gymnadenia sect. Praegnatiana Eccarius.
Gymnadenia nothosect. Praegeiniana Z. H. Feng, nothosect. nov. = Gymnadenia sect. Praegnatiana Eccarius × Gymnadenia sect. Wettsteiniana Eccarius.
Himantoglossum Spreng.
Himantoglossum nothosubgen. Comptoglossum (Karatzas) Z. H. Feng, comb. & stat. nov. = Himantoglossum subgen. Comperia (K. Koch) R. M. Bateman, Molnar & Sramkó × Himantoglossum subgen. Himantoglossum. — Basionym: × Comptoglossum Karatzas, J. Eur. Orch. 36(4): 954. 2004.
Himantoglossum nothosect. Comprinum Z. H. Feng, nothosect. nov. = Himantoglossum subgen. Comperia (K. Koch) R. M. Bateman, Molnar & Sramkó × Himantoglossum sect. Caprinum R. M. Bateman, Molnar & Sramkó.
Ophrys L.
Ophrys nothosect. Apidiniferae Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Apiflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Fuciflorae Rchb. f.
Ophrys nothosect. Apiliflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Bombyliflorae Rchb. f.
Ophrys nothosect. Apimuelleri Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Bornmuelleri Hennecke.
Ophrys nothosect. Apiniferae Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Araniferae Rchb. f.
Ophrys nothosect. Apiraniflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f.
Ophrys nothosect. Apiraniophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Ophrys.
Ophrys nothosect. Apispeculum Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Speculum Hennecke.
Ophrys nothosect. Apophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Ophrys.
Ophrys nothosect. Aracidophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Pseudophrys Godfery.
Ophrys nothosect. Araneudophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Pseudophrys Godfery.
Ophrys nothosect. Araniciflerae Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Aranispeculum Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Speculum Hennecke.
Ophrys nothosect. Aranomiflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f.
Ophrys nothosect. Aranophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Ophrys.
Ophrys nothosect. Aranyliflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Bombyliflorae Rchb. f.
Ophrys nothosect. Arelleri Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Bornmuelleri Hennecke.
Ophrys nothosect. Ariflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f.
Ophrys nothosect. Arumbilicatae Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Umbilicatae P. Quentin.
Ophrys nothosect. Bombeculum Z. H. Feng, nothosect. nov. = Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Speculum Hennecke.
Ophrys nothosect. Bomdophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Pseudophrys Godfery.
Ophrys nothosect. Bomelleri Z. H. Feng, nothosect. nov. = Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Bornmuelleri Hennecke.
Ophrys nothosect. Bominiferae Z. H. Feng, nothosect. nov. = Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Boriflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Bornmuelleri Hennecke × Ophrys sect. Fuciflorae Rchb. f.
Ophrys nothosect. Bornicatae Z. H. Feng, nothosect. nov. = Ophrys sect. Bornmuelleri Hennecke × Ophrys sect. Umbilicatae P. Quentin.
Ophrys nothosect. Faniophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Ophrys.
Ophrys nothosect. Fuceudophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Pseudophrys Godfery.
Ophrys nothosect. Fuciculum Z. H. Feng, nothosect. nov. = Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Speculum Hennecke.
Ophrys nothosect. Fuciliflorae (Z. H. Feng) Z. H. Feng, stat. nov. = Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f. — Basionym: Ophrys nothosubgen. Fuciliflorae Z. H. Feng, this publication.
Ophrys nothosect. Fuciniferae Z. H. Feng, nothosect. nov. = Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Fucophrys (Z. H. Feng) Z. H. Feng, stat. nov. = Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Ophrys. — Basionym: Ophrys nothosubgen. Fucophrys Z. H. Feng, this publication.
Ophrys nothosect. Pseudiniferae Z. H. Feng, nothosect. nov. = Ophrys sect. Pseudophrys Godfery × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Pseudolleri Z. H. Feng, nothosect. nov. = Ophrys sect. Bornmuelleri Hennecke × Ophrys sect. Pseudophrys Godfery.
Ophrys nothosect. Pseudospeculum Z. H. Feng, nothosect. nov. = Ophrys sect. Pseudophrys Godfery × Ophrys sect. Speculum Hennecke.
Ophrys nothosect. Speniferae Z. H. Feng, nothosect. nov. = Ophrys sect. Speculum Hennecke × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Spicatae Z. H. Feng, nothosect. nov. = Ophrys sect. Speculum Hennecke × Ophrys sect. Umbilicatae P. Quentin.
Ophrys nothosect. Tencatae Z. H. Feng, nothosect. nov. = Ophrys sect. Tenthrediniferae P. Quentin × Ophrys sect. Umbilicatae P. Quentin.
Ophrys nothosect. Tenthraniferae Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Umbiferae Z. H. Feng, nothosect. nov. = Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Umbilicatae P. Quentin.
Ophrys nothosect. Umbiliflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Umbilicatae P. Quentin.
Ophrys nothosubgen. Fuciliflorae Z. H. Feng, nothosubgen. nov. = Ophrys subgen. Bombyliflorae Hennecke × Ophrys subgen. Fuciflorae Hennecke & S. Munzinger.
Ophrys nothosubgen. Fucophrys Z. H. Feng, nothosubgen. nov. = Ophrys subgen. Fuciflorae Hennecke & S. Munzinger × Ophrys subgen. Ophrys.
Orchis L.
Orchis nothosect. Anthroculae Z. H. Feng, nothosect. nov. = Orchis sect. Anthropophorae H. Kretzschmar, Eccarius & H. Dietr. × Orchis sect. Masculae Lindl.
Orchis nothosect. Anthrolicae Z. H. Feng, nothosect. nov. = Orchis sect. Anthropophorae H. Kretzschmar, Eccarius & H. Dietr. × Orchis sect. Italicae H. Kretzschmar, Eccarius & H. Dietr.
Orchis nothosect. Anthrorchis Z. H. Feng, nothosect. nov. = Orchis sect. Anthropophorae H. Kretzschmar, Eccarius & H. Dietr. × Orchis sect. Orchis.
Orchis nothosect. Italorchis Z. H. Feng, nothosect. nov. = Orchis sect. Italicae H. Kretzschmar, Eccarius & H. Dietr. × Orchis sect. Orchis.
Orchis nothosect. Mascalcare Z. H. Feng, nothosect. nov. = Orchis sect. Masculae Lindl. × Orchis sect. Robustocalcare Hautz.
Orchis nothosect. Masciales Z. H. Feng, nothosect. nov. = Orchis sect. Masculae Lindl. × Orchis sect. Provinciales Parl.
Orchis nothosect. Mascillae Z. H. Feng, nothosect. nov. = Orchis sect. Masculae Lindl. × Orchis sect. Pusillae Parl.
Orchis nothosect. Mascorchis (Z. H. Feng) Z. H. Feng, stat. nov. = Orchis sect. Masculae Lindl. × Orchis sect. Orchis. — Basionym: Orchis nothosubgen. Mascorchis Z. H. Feng, this publication.
Orchis nothosect. Provalcare Z. H. Feng, nothosect. nov. = Orchis sect. Provinciales Parl. × Orchis sect. Robustocalcare Hautz.
Orchis nothosect. Provillae Z. H. Feng, nothosect. nov. = Orchis sect. Provinciales Parl. × Orchis sect. Pusillae Parl.
Orchis nothosect. Pusilcare Z. H. Feng, nothosect. nov. = Orchis sect. Pusillae Parl. × Orchis sect. Robustocalcare Hautz.
Orchis nothosubgen. Mascorchis Z. H. Feng, nothosubgen. nov. = Orchis subgen. Masculae H. Kretzschmar, Eccarius & H. Dietr. × Orchis subgen. Orchis.
Paphiopedilum Pfitzer
Paphiopedilum nothosect. Coryobata Z. H. Feng, nothosect. nov. = Paphiopedilum sect. Barbata Kraenzl. × Paphiopedilum sect. Coryopedilum Pfitzer.
Paphiopedilum nothosect. Paphiobata Z. H. Feng, nothosect. nov. = Paphiopedilum sect. Barbata Kraenzl. × Paphiopedilum sect. Paphiopedilum.
Paphiopedilum nothosect. Paphipetalum Z. H. Feng, nothosect. nov. = Paphiopedilum sect. Paphiopedilum × Paphiopedilum sect. Pardalopetalum Hallier f. & Pfitzer.
Paphiopedilum nothosect. Pardadilum Z. H. Feng, nothosect. nov. = Paphiopedilum sect. Coryopedilum Pfitzer × Paphiopedilum sect. Pardalopetalum Hallier f. & Pfitzer.
Phragmipedium Rolfe
Phragmipedium nothosect. Himantofolia Z. H. Feng, nothosect. nov. = Phragmipedium sect. Himantopetalum (Hallier f.) Garay × Phragmipedium sect. Lorifolia (Kraenzl.) Garay.
Phragmipedium nothosect. Loripedium Z. H. Feng, nothosect. nov. = Phragmipedium sect. Lorifolia (Kraenzl.) Garay × Phragmipedium sect. Phragmipedium.
Phragmipedium nothosect. Loripetalum Z. H. Feng, nothosect. nov. = Phragmipedium sect. Lorifolia (Kraenzl.) Garay × Phragmipedium sect. Platypetalum Pfitzer.
Vanda R. Br.
Vanda nothosect. Deltollaria Z. H. Feng, nothosect. nov. = Vanda sect. Deltoglossa Christenson × Vanda sect. Lamellaria Lindl.
Vanda nothosect. Longivanda Z. H. Feng, nothosect. nov. = Vanda sect. Longicalcarata Christenson × Vanda sect. Vanda.
Vanda nothosect. Testacarata Z. H. Feng, nothosect. nov. = Vanda sect. Longicalcarata Christenson × Vanda sect. Testaceae L. M. Gardiner.
Papaveraceae
Corydalis DC.
Corydalis nothosect. Corydacava Z. H. Feng, nothosect. nov. = Corydalis sect. Corydalis × Corydalis sect. Radix-cava Irmisch.
Meconopsis Vig.
Meconopsis nothosect. Granopsis Z. H. Feng, nothosect. nov. = Meconopsis sect. Grandes Fedde × Meconopsis sect. Meconopsis.
Papaver L.
Papaver nothosect. Meconantha Z. H. Feng, nothosect. nov. = Papaver sect. Macrantha Elkan × Papaver sect. Meconidium Spach.
Papaver nothosect. Papadium Z. H. Feng, nothosect. nov. = Papaver sect. Papaver × Papaver sect. Rhoeadium Spach.
Plantaginaceae
Antirrhinum L.
Antirrhinum nothosect. Streptorrhinum Z. H. Feng, nothosect. nov. = Antirrhinum sect. Antirrhinum × Antirrhinum sect. Streptosepalum (Rothm.) Fern. Casas.
Plantago L.
Plantago nothosect. Lamprotago Z. H. Feng, nothosect. nov. = Plantago sect. Lamprosantha Decne. × Plantago sect. Plantago.
Plantago nothosect. Lanceisantha Z. H. Feng, nothosect. nov. = Plantago sect. Lamprosantha Decne. × Plantago sect. Lanceifolia Barnéoud.
Plantago nothosubgen. Psyllago Z. H. Feng, nothosubgen. nov. = Plantago subgen. Plantago × Plantago subgen. Psyllium (Juss.) Harms & Reiche.
Plumbaginaceae
Limonium Mill.
Limonium nothosect. Ironium Z. H. Feng, nothosect. nov. = Limonium sect. Iranolimon M. Malekm., Akhani & Borsch × Limonium sect. Limonium.
Ranunculaceae
Helleborus L.
Helleborus nothosect. Chenoborus Z. H. Feng, nothosect. nov. = Helleborus sect. Chenopus Schiffn. × Helleborus sect. Helleborus.
Helleborus nothosect. Dicastrum Z. H. Feng, nothosect. nov. = Helleborus sect. Dicarpon Ulbr. × Helleborus sect. Helleborastrum Spach.
Helleborus nothosect. Grirus Z. H. Feng, nothosect. nov. = Helleborus sect. Griphopus Spach × Helleborus sect. Helleborus.
Helleborus nothosect. Hellarpon Z. H. Feng, nothosect. nov. = Helleborus sect. Dicarpon Ulbr. × Helleborus sect. Helleborus.
Helleborus nothosect. Hellastrum (Z. H. Feng) Z. H. Feng, stat. nov. = Helleborus sect. Helleborastrum Spach × Helleborus sect. Helleborus. — Basionym: Helleborus nothosubgen. Hellastrum Z. H. Feng, this publication.
Helleborus nothosect. Hellecarpus Z. H. Feng, nothosect. nov. = Helleborus sect. Helleborus × Helleborus sect. Syncarpus Schiffn.
Helleborus nothosubgen. Hellastrum Z. H. Feng, nothosubgen. nov. = Helleborus subgen. Helleborastrum (Spach) K. Werner & F. Ebel × Helleborus subgen. Helleborus.
Pulsatilla Mill.
Pulsatilla nothosect. Preotilla (Z. H. Feng) Z. H. Feng, stat. nov. = Pulsatilla sect. Preonanthus (DC.) Spach. × Pulsatilla sect. Pulsatilla. — Basionym: Pulsatilla nothosubgen. Preotilla Z. H. Feng, this publication.
Pulsatilla nothosect. Semipulsatilla Z. H. Feng, nothosect. nov. = Pulsatilla sect. Pulsatilla × Pulsatilla sect. Semicampanaria Zämelis & Paegle.
Pulsatilla nothoser. Patanthus Z. H. Feng, nothoser. nov. = Pulsatilla ser. Patentes (Aichele & Schweg.) Juz. ex Tamura × Pulsatilla sect. Preonanthus (DC.) Spach.
Pulsatilla nothoser. Patensatilla Z. H. Feng, nothoser. nov. = Pulsatilla ser. Patentes (Aichele & Schweg.) Juz. ex Tamura × Pulsatilla ser. Pulsatilla.
Pulsatilla nothoser. Patenses Z. H. Feng, nothoser. nov. = Pulsatilla ser. Patentes (Aichele & Schweg.) Juz. ex Tamura × Pulsatilla ser. Pratenses (Aichele & Schweg.) Juz. ex Tamura.
Pulsatilla nothoser. Prananthus Z. H. Feng, nothoser. nov. = Pulsatilla ser. Pratenses (Aichele & Schweg.) Juz. ex Tamura × Pulsatilla sect. Preonanthus (DC.) Spach.
Pulsatilla nothoser. Pratensatilla Z. H. Feng, nothoser. nov. = Pulsatilla ser. Pratenses (Aichele & Schweg.) Juz. ex Tamura × Pulsatilla ser. Pulsatilla.
Pulsatilla nothoser. Preotilla (Z. H. Feng) Z. H. Feng, stat. nov. = Pulsatilla sect. Preonanthus (DC.) Spach. × Pulsatilla ser. Pulsatilla. — Basionym: Pulsatilla nothosubgen. Preotilla Z. H. Feng, this publication.
Pulsatilla nothosubgen. Preotilla Z. H. Feng, nothosubgen. nov. = Pulsatilla subgen. Preonanthus (DC.) Juz. × Pulsatilla subgen. Pulsatilla.
Rosaceae
Cliffortia L.
Cliffortia nothosect. Acetae Z. H. Feng, nothosect. nov. = Cliffortia sect. Acerosae C. Whitehouse × Cliffortia sect. Costatae Weim.
Cliffortia nothosect. Multifiviae Z. H. Feng, nothosect. nov. = Cliffortia sect. Multifidae C. Whitehouse × Cliffortia sect. Multinerviae DC.
Cliffortia nothosect. Multoides Z. H. Feng, nothosect. nov. = Cliffortia sect. Multinerviae DC. × Cliffortia sect. Ruboides C. Whitehouse.
Cliffortia nothosect. Rufidae Z. H. Feng, nothosect. nov. = Cliffortia sect. Multifidae C. Whitehouse × Cliffortia sect. Ruboides C. Whitehouse.
Cotoneaster Medik.
Cotoneaster nothosect. Chaenaster (Z. H. Feng) Z. H. Feng, stat. nov. = Cotoneaster sect. Chaenopetalum Koehne × Cotoneaster sect. Cotoneaster. — Basionym: Cotoneaster nothosubgen. Chaenaster Z. H. Feng, this publication.
Cotoneaster nothosect. Chaenoflori Z. H. Feng, nothosect. nov. = Cotoneaster sect. Chaenopetalum Koehne × Cotoneaster sect. Densiflori T. T. Yu.
Cotoneaster nothoser. Frinosi Z. H. Feng, nothoser. nov. = Cotoneaster ser. Frigidi T. T. Yu × Cotoneaster ser. Pannosi Flinck & B. Hylmö.
Cotoneaster nothoser. Concantes Z. H. Feng, nothoser. nov. = Cotoneaster ser. Conspicui G. Klotz × Cotoneaster ser. Radicantes G. Klotz.
Cotoneaster nothoser. Frifolii Z. H. Feng, nothoser. nov. = Cotoneaster ser. Frigidi T. T. Yu × Cotoneaster ser. Salicifolii T. T. Yu.
Cotoneaster nothoser. Racemaster Z. H. Feng, nothoser. nov. = Cotoneaster ser. Cotoneaster × Cotoneaster ser. Racemiflori (Pojark.) G. Klotz.
Cotoneaster nothoser. Tomentaster Z. H. Feng, nothoser. nov. = Cotoneaster ser. Cotoneaster × Cotoneaster ser. Tomentosi J. Fryer & B. Hylmö.
Cotoneaster nothosubgen. Chaenaster Z. H. Feng, nothosubgen. nov. = Cotoneaster subgen. Chaenopetalum (Koehne) G. Klotz × Cotoneaster subgen. Cotoneaster.
Crataegus L.
Crataegus nothosect. Brespilus (Z. H. Feng) Z. H. Feng, stat. nov. = Crataegus sect. Brevispinae (Beadle) C. K. Schneid. × Crataegus sect. Mespilus (L.) T. A. Dickinson & E. Y. Y. Lo. — Basionym: Crataegus nothosubgen. Brespilus Z. H. Feng, this publication.
Crataegus nothosect. Coccanthae Z. H. Feng, nothosect. nov. = Crataegus sect. Coccineae Loudon × Crataegus sect. Macracanthae Loudon.
Crataegus nothoser. Anofoliae Z. H. Feng, nothoser. nov. = Crataegus ser. Anomalae (Sarg. ex Eggl.) Rehder × Crataegus ser. Tenuifoliae (Sarg. ex Eggl.) Rehder.
Crataegus nothoser. Anosae Z. H. Feng, nothoser. nov. = Crataegus ser. Anomalae (Sarg. ex Eggl.) Rehder × Crataegus ser. Pruinosae Sarg. ex Rehder.
Crataegus nothoser. Anothae Z. H. Feng, nothoser. nov. = Crataegus ser. Anomalae (Sarg. ex Eggl.) Rehder × Crataegus ser. Macracanthae (Loudon) Rehder.
Crataegus nothoser. Apolles Z. H. Feng, nothoser. nov. = Crataegus ser. Apiifoliae (Loudon) Rehder × Crataegus ser. Molles (Sarg. ex C. K. Schneider) Rehder.
Crataegus nothoser. Coccifoliae Z. H. Feng, nothoser. nov. = Crataegus ser. Coccineae (Loudon) Rehder × Crataegus ser. Tenuifoliae (Sarg. ex Eggl.) Rehder.
Crataegus nothoser. Crataegrae Z. H. Feng, nothoser. nov. = Crataegus ser. Crataegus × Crataegus ser. Nigrae (Loudon) Russanov.
Crataegus nothoser. Crusanthae Z. H. Feng, nothoser. nov. = Crataegus ser. Crus-galli (Loudon) Rehder × Crataegus ser. Macracanthae (Loudon) Rehder.
Crataegus nothoser. Incanthae Z. H. Feng, nothoser. nov. = Crataegus ser. Intricatae Sarg. ex Rehder × Crataegus ser. Macracanthae (Loudon) Rehder.
Crataegus nothoser. Innosae Z. H. Feng, nothoser. nov. = Crataegus ser. Intricatae Sarg. ex Rehder × Crataegus ser. Pruinosae Sarg. ex Rehder.
Crataegus nothoser. Macoliae Z. H. Feng, nothoser. nov. = Crataegus ser. Macracanthae (Loudon) Rehder × Crataegus ser. Parvifoliae (Loudon) Rehder.
Crataegus nothoser. Manosae Z. H. Feng, nothoser. nov. = Crataegus ser. Macracanthae (Loudon) Rehder × Crataegus ser. Pruinosae Sarg. ex Rehder.
Crataegus nothoser. Mollides Z. H. Feng, nothoser. nov. = Crataegus ser. Molles (Sarg. ex C. K. Schneider) Rehder × Crataegus ser. Virides (Beadle ex C. K. Schneider) Rehder.
Crataegus nothoser. Potigynae Z. H. Feng, nothoser. nov. = Crataegus ser. Pentagynae (C. K. Schneid.) Russanov × Crataegus ser. Poticae Pojark.
Crataegus nothoser. Prudifoliae Z. H. Feng, nothoser. nov. = Crataegus ser. Pruinosae Sarg. ex Rehder × Crataegus ser. Rotundifoliae (Eggl. ex Eggl.) Rehder.
Crataegus nothoser. Pugalli Z. H. Feng, nothoser. nov. = Crataegus ser. Crus-galli (Loudon) Rehder × Crataegus ser. Punctatae (Loudon) Rehder.
Crataegus nothoser. Puncosae Z. H. Feng, nothoser. nov. = Crataegus ser. Pruinosae Sarg. ex Rehder × Crataegus ser. Punctatae (Loudon) Rehder.
Crataegus nothoser. Rocanthae Z. H. Feng, nothoser. nov. = Crataegus ser. Rotundifoliae (Eggl. ex Eggl.) Rehder × Crataegus ser. Macracanthae (Loudon) Rehder.
Crataegus nothoser. Ronuifoliae Z. H. Feng, nothoser. nov. = Crataegus ser. Rotundifoliae (Eggl. ex Eggl.) Rehder × Crataegus ser. Tenuifoliae (Sarg.) Rehder.
Crataegus nothoser. Tentae Z. H. Feng, nothoser. nov. = Crataegus ser. Punctatae (Loudon) Rehder × Crataegus ser. Tenuifoliae (Sarg. ex Eggl.) Rehder.
Crataegus nothosubgen. Ametaegus Z. H. Feng, nothosubgen. nov. = Crataegus subgen. Americanae El Gazzar × Crataegus subgen. Crataegus.
Crataegus nothosubgen. Brespilus Z. H. Feng, nothosubgen. nov. = Crataegus subgen. Brevispinae (Beadle) Ufimov & T. A. Dickinson × Crataegus subgen. Mespilus (L.) Ufimov & T. A. Dickinson.
Crataegus nothosubgen. Crataeguineae (K. I. Chr.) Z. H. Feng, stat. nov. = Crataegus subgen. Crataegus × Crataegus subgen. Sanguineae Ufimov. — Basionym: Crataegus nothosect. Crataeguineae K. I. Chr., Syst. Bot. Monogr. 35: 161. 1992.
Prunus L.
Prunus nothosect. Micropurus Z. H. Feng, nothosect. nov. = Prunus sect. Microcerasus (Spach) C. K. Schneid. × Prunus sect. Prunus.
Prunus nothosect. Prudalus Z. H. Feng, nothosect. nov. = Prunus sect. Amygdalus (L.) Benth. & Hook. f. × Prunus sect. Prunus.
Prunus nothosect. Pruniaca Z. H. Feng, nothosect. nov. = Prunus sect. Armeniaca (Scop.) Koch × Prunus sect. Prunus.
Pyrus L.
Pyrus nothosubgen. Pyshia Z. H. Feng, nothosubgen. nov. = Pyrus subgen. Pashia (Koehne) B. B. Liu × Pyrus subgen. Pyrus.
Rubus L.
Rubus nothosubgen. Bachobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Batothamnus (Focke) E. H. L. Krause × Rubus subgen. Malachobatus (Focke) Fritsch.
Rubus nothosubgen. Bateobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Batothamnus (Focke) E. H. L. Krause × Rubus subgen. Idaeobatus Focke.
Rubus nothosubgen. Chalanobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Chamaerubus Kuntze × Rubus subgen. Melanobatus (Greene) House.
Rubus nothosubgen. Cynobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Cylactis (Raf.) Focke × Rubus subgen. Melanobatus (Greene) House.
Rubus nothosubgen. Idanobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Idaeobatus Focke × Rubus subgen. Melanobatus (Greene) House.
Rubus nothosubgen. Linchobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Lineati (Focke) T. R. Huang & X. Y. Zhu × Rubus subgen. Malachobatus (Focke) Fritsch.
Rubus nothosubgen. Malnobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Malachobatus (Focke) Fritsch × Rubus subgen. Melanobatus (Greene) House.
Rubus nothosubgen. Rubobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Melanobatus (Greene) House × Rubus subgen. Rubus.
Saxifragaceae
Saxifraga L.
Saxifraga nothosect. Cotypera Z. H. Feng, nothosect. nov. = Saxifraga sect. Cotylea Tausch × Saxifraga sect. Gymnopera D. Don.
Saxifraga nothosect. Liguphyrion Z. H. Feng, nothosect. nov. = Saxifraga sect. Ligulatae Haw. × Saxifraga sect. Porphyrion Tausch.
Saxifraga nothosect. Ligusaxifraga Z. H. Feng, nothosect. nov. = Saxifraga sect. Ligulatae Haw. × Saxifraga sect. Saxifraga.
Saxifraga nothosect. Trachysaxifraga Z. H. Feng, nothosect. nov. = Saxifraga sect. Saxifraga × Saxifraga sect. Trachyphyllum (Gaudin) W. D. J. Koch.
Saxifraga nothosubsect. Androsaxifraga Z. H. Feng, nothosubsect. nov. = Saxifraga subsect. Androsaceae (Engl. & Irmsch.) Tkach, Röser & M. H. Hoffm. × Saxifraga subsect. Saxifraga.
Saxifraga nothosubsect. Androtylites Z. H. Feng, nothosubsect. nov. = Saxifraga subsect. Androsaceae (Engl. & Irmsch.) Tkach, Röser & M. H. Hoffm. × Saxifraga subsect. Tridactylites (Haw.) Gornall.
Saxifraga nothosubsect. Araxifraga Z. H. Feng, nothosubsect. nov. = Saxifraga subsect. Arachnoideae (Engl. & Irmsch.) Tkach, Röser & M. H. Hoffm. × Saxifraga subsect. Saxifraga.
Saxifraga nothosubsect. Kabrrosae Z. H. Feng, nothosubsect. nov. = Saxifraga subsect. Kabschia Rouy & E. G. Camus × Saxifraga subsect. Squarrosae (Engl. & Irmsch.) Tkach, Röser & M. H. Hoffm.
Saxifraga nothosubsect. Kabulatae Z. H. Feng, nothosubsect. nov. = Saxifraga sect. Ligulatae Haw. × Saxifraga subsect. Kabschia Rouy & E. G. Camus.
Saxifraga nothosubsect. Ligusaxifraga (Z. H. Feng) Z. H. Feng, stat. nov. = Saxifraga sect. Ligulatae Haw. × Saxifraga subsect. Saxifraga. — Basionym: Saxifraga nothosect. Ligusaxifraga Z. H. Feng, this publication.
Saxifraga nothosubsect. Murrosae Z. H. Feng, nothosubsect. nov. = Saxifraga subsect. Mutatae (Engl. & Irmsch.) Gornall × Saxifraga subsect. Squarrosae (Engl. & Irmsch.) Tkach, Röser & M. H. Hoffm.
Saxifraga nothosubsect. Mutalatae Z. H. Feng, nothosubsect. nov. = Saxifraga sect. Ligulatae Haw. × Saxifraga subsect. Mutatae (Engl. & Irmsch.) Gornall.
Saxifraga nothosubsect. Trachysaxifraga (Z. H. Feng) Z. H. Feng, stat. nov. = Saxifraga subsect. Saxifraga × Saxifraga sect. Trachyphyllum (Gaudin) W. D. J. Koch. — Basionym: Saxifraga nothosect. Trachysaxifraga Z. H. Feng, this publication.
Saxifraga nothosubsect. Trisaxifraga Z. H. Feng, nothosubsect. nov. = Saxifraga subsect. Saxifraga × Saxifraga subsect. Tridactylites (Haw.) Gornall.
Verbenaceae
Verbena L.
Verbena nothoser. Bracelabrae Z. H. Feng, nothoser. nov. = Verbena ser. Bracteatae G. L. Nesom × Verbena ser. Candelabrae G. L. Nesom.
Verbena nothoser. Canbena Z. H. Feng, nothoser. nov. = Verbena ser. Candelabrae G. L. Nesom × Verbena ser. Verbena.
Verbena nothoser. Candachyae Z. H. Feng, nothoser. nov. = Verbena ser. Candelabrae G. L. Nesom × Verbena ser. Leptostachyae Schauer.
Verbena nothoser. Candelices Z. H. Feng, nothoser. nov. = Verbena ser. Candelabrae G. L. Nesom × Verbena ser. Simplices G. L. Nesom.
Verbena nothoser. Halabrae Z. H. Feng, nothoser. nov. = Verbena ser. Candelabrae G. L. Nesom × Verbena ser. Haleae G. L. Nesom.
Verbena nothoser. Leptacteatae Z. H. Feng, nothoser. nov. = Verbena ser. Bracteatae G. L. Nesom × Verbena ser. Leptostachyae Schauer.
Verbena nothoser. Leptices Z. H. Feng, nothoser. nov. = Verbena ser. Leptostachyae Schauer × Verbena ser. Simplices G. L. Nesom.
Verbena nothoser. Plibena Z. H. Feng, nothoser. nov. = Verbena ser. Plicatae G. L. Nesom × Verbena ser. Verbena.
Verbena nothoser. Plicteatae Z. H. Feng, nothoser. nov. = Verbena ser. Bracteatae G. L. Nesom × Verbena ser. Plicatae G. L. Nesom.
Violaceae
Viola L.
Viola nothosect. Plagioviola Z. H. Feng, nothosect. nov. = Viola sect. Plagiostigma Godr. × Viola sect. Viola.
Viola nothosubsect. Brebracteatae Z. H. Feng, nothosubsect. nov. = Viola subsect. Bracteolatae Kupffer × Viola subsect. Ebracteatae Kupffer.
Viola nothosubsect. Rostellares Z. H. Feng, nothosubsect. nov. = Viola subsect. Patellares (Boiss.) Rouy & Foucaud × Viola subsect. Rostratae (Kupffer) W. Becker.
Viola nothosubsect. Rostiola Z. H. Feng, nothosubsect. nov. = Viola subsect. Rostratae (Kupffer) W. Becker × Viola subsect. Viola.
3.3. New Nothogenera and Combinations of Pteridophytes and Spermatophytes
Apocynaceae
× Orbelia plowesii (P. V. Heath) Z. H. Feng, comb. nov. — Basionym: × Gonostapelia plowesii P. V. Heath, Calyx 1(1): 21. 1992.
Hybrid formula: Orbea maculata (N. E. Br.) L. C. Leach × Stapelia kwebensis N. E. Br.
Type: Zimbabwe. Mutema Pan, s. d., Plowes 3798 (holotype: PRE; possible isotype: SRGH).
× Tromostapelia bella (A. Berger) Z. H. Feng, comb. nov. — Basionym: Stapelia × bella A. Berger, Gard. Chron. ser. 3, 31: 137. 1902.
Hybrid formula: Stapelia deflexa Jacq. × Tromotriche revoluta (Masson) Haw.
Type: Provenance not given, 16 October 1901, A. Berger s. n. (holotype: K [barcode] K 000306220, digital image!).
Athyriaceae
Cornopteris bipinnata (K. Hori) Z. H. Feng, comb. nov. — Basionym: Athyrium bipinnatum K. Hori, PhytoKeys 148: 97. 2020.
Type: JAPAN. Shikoku: Kochi prefecture, Ochi town, Mt. Yokogura, 664 m, planted coniferous forest containing Cryptomeria japonica (Thunb. ex L. f.) D. Don, on soil, 29 June 2019, K. Hori 3277 (holotype: MAK [barcode] MAK 466762; isotype: MBK).
Cornopteris × decurrentiserrata (K. Hori) Z. H. Feng, comb. nov. — Basionym: Athyrium × decurrentiserratum K. Hori, Fern Gaz. 21(6): 262. 2021.
Hybrid formula: Cornopteris crenulatoserrulata (Makino) Nakai × Cornopteris decurrentialata (Hook.) Nakai.
Type: JAPAN. Honshu: Tokyo-to, Ome city, Koshizawa, 396 m, planted coniferous forest containing Cryptomeria japonica (Thunb. ex L. f.) D. Don, on soil, 4 June 2018, Hori 2981 (holotype: MAK [barcode] MAK 466750; isotype: MBK).
Cornopteris × pseudobipinnata (K. Hori) Z. H. Feng, comb. nov. — Basionym: Athyrium × pseudobipinnatum K. Hori, Fern Gaz. 21(6): 264. 2021.
Hybrid formula: Cornopteris bipinnata (K. Hori) Z. H. Feng × Cornopteris decurrentialata (Hook.) Nakai.
Type: JAPAN. Kyushu: Fukuoka pref., Itoshima city, Shiraito, 515 m, planted coniferous forest containing Cryptomeria japonica, on soil, 23 June 2019, Hori 3274 (holotype: MAK [barcode] MAK 466761; isotype: MBK).
Cornopteris × subchristenseniana (K. Hori) Z. H. Feng, comb. nov. — Basionym: Athyrium × subchristensenianum K. Hori, Fern Gaz. 21(6): 265. 2021.
Hybrid formula: Cornopteris crenulatoserrulata (Makino) Nakai × an unknown allotetraploid species.
Type: JAPAN. Honshu: Akita pref., Yurihonjo city, Hachinosawa, 180 m, planted coniferous forest containing Cryptomeria japonica (Thunb. ex L. f.) D. Don, on soil, 5 July 2019, Hori 3281 (holotype: MAK [barcode] MAK 466765; isotype: MBK).
Convolvulaceae
Convolvulus × turcicus nothosubsp. peshmenii (Aykurt & Sümbül) Z. H. Feng, comb. & stat. nov. — Basionym: Convolvulus × peshmenii Aykurt & Sümbül, Nordic J. Bot. 29(4): 409. 2011.
Hybrid formula: Convolvulus boissieri subsp. compactus (Boiss.) Stace × Convolvulus holosericeus subsp. macrocalycinus Hausskn. & Bornm.
Type: Turkey. southern Anatolia. Adana, Pozantı, 5-6 km from Pozantı to Adana, 945 m, 9 June 2007, C. Aykurt & N. Kemaloğlu 1495 (holotype: AKDU).
Lamiaceae
Salvia × hegelmaieri nothosubsp. accidentalis (Sánchez-Gómez & R. Morales) Z. H. Feng, comb. & stat. nov. — Basionym: Salvia × accidentalis Sánchez-Gómez & R. Morales, Anales Jard. Bot. Madrid 57(2): 421. 2000.
Hybrid formula: Salvia blancoana subsp. vellerea (Cuatrec.) W. Lippert × Salvia officinalis subsp. officinalis.
Type: SPAIN. Murcia. Moratalla, La Alberquilla, 10 May 1993, P. Sánchez Gómez s. n. (holotype: MA [barcode] MA 593325).
Orchidaceae
Calanthe × berryana (Sander) Z. H. Feng, comb. nov. — Basionym: × Phaiocalanthe berryana Sander, Gard. Chron. ser. 3, 18: 655. 1895.
Hybrid formula: Calanthe humblotii (Rchb. f.) M. W. Chase, Christenh. & Schuit. × Calanthe masuca (D. Don) Lindl.
Type: Not designated.
Calanthe × brandtiae (Sander) Z. H. Feng, comb. nov. — Basionym: × Phaiocalanthe brandtiae Sander, Gard. Chron. ser. 3, 21: 115. 1897.
Hybrid formula: Calanthe tankervilleae (Banks) M. W. Chase, Christenh. & Schuit. × [Calanthe × veitchii Paxton & Lindl.].
Type: Not designated.
Calanthe × inspirata (Veitch) Z. H. Feng, comb. nov. — Basionym: × Phaiocalanthe inspirata Veitch, Gard. Chron. ser. 3, 22: 315. 1897.
Hybrid formula: Calanthe masuca (D. Don) Lindl. × Calanthe tankervilleae (Banks) M. W. Chase, Christenh. & Schuit.
Type: Not designated.
Poaceae
Anthoxanthum × zinserlingii (Tzvelev) Z. H. Feng, comb. nov. — Basionym: Hierochloe × zinserlingii Tzvelev, Novosti Sist. Vyssh. Rast. 10: 83. 1973.
Hybrid formula: Anthoxanthum monticola (Bigelow) Veldkamp × Anthoxanthum nitens (Weber) Y. Schouten & Veldkamp.
Type: RUSSIA. Ural. on the cliffs of Mount Sablja, 20 July 1926, G. Zinserling 504 (holotype: LE [barcode] LE 01078250, digital image!).
× Pseudelymus versicolor (A. P. Khokhr.) Z. H. Feng, comb. nov. — Basionym: Elymus × versicolor A. P. Khokhr., Biol. Rast. Fl. Sev. Dal'n. Vostok.: 13. 1981.
Hybrid formula: Elymus macrourus (Turcz. ex Steud.) Tzvelev × Pseudoroegneria gmelinii (Trin. ex Schrad.) Sennikov.
Type: RUSSIA. Magadan Region. Srednekansky District, mountain Zamkovyi, mosses by the rock, 22 August 1979, S. Ershova, P. Khokryakov, A. P. Khokryakov s. n. (holotype: MHA [barcode] MHA 0359393, digital image!; isotypes: MAG [barcodes] MAG 0000235-MAG 0000241, digital images!; VLA [barcode] VLA 00001831).
Rosaceae
× Arsorbus abscondita (Kovanda) Z. H. Feng, comb. nov. — Basionym: Sorbus × abscondita Kovanda, Verh. Zool.-Bot. Ges. Österreich 133: 339. 1996.
Hybrid formula: Aria danubialis (Jáv.) Sennikov & Kurtto × Sorbus aucuparia L.
Type: Czech Republic. Bohemia occid.-centr.: in angulo clivi Pochvalovská stráň dicti situ septentr.-orient. a pago Pochvalov (distr. Louny); solo calcifero-schistaceo, alt. 490 m, 25 May 1992, M. Kovanda s. n. (holotype: WU; isotypes: PR).
Eriolobus angustifolius (Aiton) Su Liu ex Z. H. Feng, comb. nov. — Basionym: Pyrus angustifolia Aiton, Hort. Kew. [W. Aiton] 2: 176. 1789.
Neotype (designated by Li et al. : 71): USA. North Carolina, locality unknown, s. d., Greene s. n. (neotype: K [barcode] K 000758465, digital image!).
Eriolobus coronarius (L.) Su Liu ex Z. H. Feng, comb. nov. — Basionym: Pyrus coronaria L., Sp. Pl. [Linnaeus] 1: 480. 1753.
Type: Provenance not given. s. d., Linnaeus s. n., Herb. Linn. No. 647.5 (holotype: LINN [barcode] LINN 647.5, digital image!).
Eriolobus ioensis (Alph. Wood) Su Liu ex Z. H. Feng, comb. & stat. nov. — Basionym: Pyrus coronaria var. ioensis Alph. Wood, Class-book Bot. (ed. 1861): 333. 1861.
Neotype (designated by Langenfeld : 153): USA. Missouri, Daviess, near Pattonsburg, thickets, limestone uplands, 14 June 1924, E. J. Palmer 25466 (neotype: S).
× Eriomalus Su Liu ex Z. H. Feng, nothogen. nov.
Hybrid formula: Eriolobus (DC.) M. Roem. × Malus Mill.
× Eriomalus heterophylla (Spach) Z. H. Feng, comb. nov. — Basionym: Malus × heterophylla Spach, Hist. Nat. Vég. (Spach) 2: 138. 1834, pro sp.
Hybrid formula: Eriolobus coronarius (L.) Su Liu ex Z. H. Feng × Malus domestica (Suckow) Borkh.
Type: Not designated.
× Eriomalus soulardii (L. H. Bailey) Su Liu ex Z. H. Feng, comb. nov. — Basionym: Pyrus × soulardii L. H. Bailey (‘soulardi’), Amer. Garden 12: 472. 1891, pro sp.
Hybrid formula: Eriolobus ioensis (Alph. Wood) Su Liu ex Z. H. Feng × Malus domestica (Suckow) Borkh.
Type: Not designated.
× Tormariosorbus liljeforsii (T. C. G. Rich) Z. H. Feng, comb. nov. — Basionym: Sorbus × liljeforsii T. C. G. Rich, Nordic J. Bot. 25(5-6): 339. 2008.
Hybrid formula: Sorbus aucuparia L. × [× Tormariosorbus intermedia (Ehrh.) J. M. H. Shaw].
Type: Sweden. Västergötland: Finnerödja, Brinken, 10 July 1920, J. A. O. Skårman s. n. (holotype: UPS).
Saxifragaceae
Micranthes × crawfordii (E. S. Marshall) Z. H. Feng, comb. nov. — Basionym: Saxifraga × crawfordii E. S. Marshall, J. Bot. 47: 98. 1938.
Hybrid formula: Micranthes nivalis (L.) Small × Micranthes stellaris (L.) Galasso, Banfi & Soldano.
Type: Not designated.
Thelypteridaceae
× Abacopterella jerdonii (Ching) Z. H. Feng, comb. nov. — Basionym: Cyclosorus jerdonii Ching, Bull. Fan Mem. Inst. Biol. Bot. 8: 228. 1938.
Hybrid formula: Abacopteris nudata (Roxb.) S. E. Fawc. & A. R. Sm. × Christella procera (D. Don) Mazumdar.
Type: SIKKIM [India]. Jerdon s. n. (holotype: K [barcode] K 000951517, digital image!; isotypes: B [barcodes] B 200041322, B 200041323, B 200041324, digital images!).
× Chririx Z. H. Feng, nothogen. nov.
Hybrid formula: Christella H. Lév. × Grypothrix (Holttum) S. E. Fawc. & A. R. Sm.
× Chririx gogoii (Fraser-Jenk.) Z. H. Feng, comb. nov. — Basionym: Thelypteris × gogoii Fraser-Jenk., New Sp. Syndr. Indian Pteridol.: 248. 1997.
Hybrid formula: Christella procera (D. Don) Mazumdar × Grypothrix triphylla (Sw.) S. E. Fawc. & A. R. Sm.
Type: INDIA. Assam, Jorhat District, Goiaghat, shortly north of Kosarihat village, 21 December 1995, C. R Fraser-Jenkins 23846, FN 3001 (holotype: BM).
× Chririx thwaitesii (Hook.) Z. H. Feng, comb. nov. — Basionym: Meniscium × thwaitesii Hook., Fil. Exot. 2: t. 83. 1858.
Hybrid formula: Christella parasitica (L.) H. Lév. × Grypothrix triphylla (Sw.) S. E. Fawc. & A. R. Sm.
Type: SRI LANKA. s. d., G. H. K. Thwaites 3145 (holotype: K [barcode] K 000951504, digital image!; isotypes: B [barcodes] B 200068843, B 200068848, BM [barcode] BM 001044868, GH [barcode] GH 00112616, K [barcode] K 000951505, digital image!, L [barcode] L 0061084, P [barcodes] P 00643850, P 00643851, UC [barcode] UC 267428).
× Chrismatopteris varievenulosa (Viane) Z. H. Feng, comb. nov. — Basionym: Thelypteris × varievenulosa Viane, Bull. Soc. Roy. Bot. Belgique 118(1): 49. 1985.
Hybrid formula: Christella hispidula (Decne.) Holttum × Pneumatopteris afra (Christ) Holttum.
Type: WEST AFRICA [IVORY COAST]. Mt. Tonkoui. Wet, shaded trackside at ca. 900 m alt., among Cyathea camerooniana and Thelypteris afra, 29 July 1979, R. Viane 1030 (holotype: GENT [barcode] GENT 0000090035227, digital image!; isotype: GENT [barcode] GENT 0000090035166, digital image!).
× Christelliopsis nepalensis (Fraser-Jenk.) Z. H. Feng, comb. nov. — Basionym: Thelypteris × nepalensis Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes: 188. 2008.
Hybrid formula: Christella dentata (Forssk.) Brownsey & Jermy × Menisciopsis penangiana (Hook.) S. E. Fawc. & A. R. Sm.
Type: NEPAL. Nawalparasi District, Lumbini Zone, c. 1 km northeast of Daunne near top of pass over Siwalik ridge, shortly south of Dumkibas, east of Butwal on road to Narayanghat, 28 November 2005, C. R. Fraser-Jenkins 31892 (holotype: TAIF [barcode] TAIF 309339, digital image!).
× Glaphyronisciopsis Z. H. Feng, nothogen. nov.
Hybrid formula: Glaphyropteridopsis Ching × Menisciopsis (Holttum) S. E. Fawc. & A. R. Sm.
× Glaphyronisciopsis occulta (C. Hope) Z. H. Feng, comb. nov. — Basionym: Nephrodium occultum C. Hope, J. Bombay Nat. Hist. Soc. 12: 627, t. 13. 1899.
Hybrid formula: Glaphyropteridopsis erubescens (Wall. ex Hook.) Ching × Menisciopsis penangiana (Hook.) S. E. Fawc. & A. R. Sm.
Lectotype (designated by Singh and Rawat : 40): INDIA. North-western India, Uttarakhand, 1890, P. W. Mackinnon & V. A. Mackinnon s. n. (lectotype: BM [barcode] BM 001044870, digital image!).
Grypothrix × pseudoliukiuensis (Seriz.) Z. H. Feng, comb. nov. — Basionym: Thelypteris × pseudoliukiuensis Seriz., J. Phytogeogr. Taxon. 29(1): 25. 1981.
Hybrid formula: Grypothrix cuspidata (Blume) S. E. Fawc. & A. R. Sm. × Grypothrix triphylla (Sw.) S. E. Fawc. & A. R. Sm.
Type: RYUKYU ISLAND. Mt. Komidake, alt. ca. 90 m, 9 August 1971, S. Serizawa 14678 (holotype: AICH).
4. Conclusion
Accurate taxonomy is not merely an academic exercise but a vital tool for safeguarding biodiversity. By refining classifications and elucidating hybrid dynamics, the current work presents a robust update to the taxonomy and nomenclature of selected vascular plant genera, introducing critical nomenclatural novelties, including revised infrageneric classifications and new nothoranks. The updated classifications demonstrate the necessity of reconciling molecular data with morphological traits to resolve long-standing taxonomic ambiguities, and the validation of hybrid nothoranks addresses gaps in hybrid nomenclature, providing a structured approach to categorize interspecific and intergeneric hybrids. As phylogenetic data continue to accumulate, the taxonomic community must remain agile, embracing iterative revisions that reflect the living, evolving nature of the plant kingdom. Such efforts ensure that our scientific frameworks remain robust, relevant, and responsive to the dual challenges of biodiversity loss and environmental change.
Abbreviations

APG

Angiosperm Phylogeny Group

ICN

International Code of Nomenclature for Algae, Fungi, and Plants

IPNI

The International Plant Names Index

PPG

Pteridophyte Phylogeny Group

Acknowledgments
I am deeply indebted to librarians from the library of Kew for the interlibrary loan (ILL) service. My thanks go to Ms. Ying-ying Wu for polishing language. I thank colleagues of BAZI for critically reviewing the manuscript before submission and for keen comments and advice from anonymous referee(s) that contribute to the final version of the manuscript.
Author Contributions
Zhen-Hao Feng is the sole author. The author read and approved the final manuscript.
Conflicts of Interest
The author declares no conflicts of interest.
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    Feng, Z. (2025). New Ranks and New Combinations in Selected Families and Genera. Plant, 13(2), 76-99. https://doi.org/10.11648/j.plant.20251302.14

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    Feng, Z. New Ranks and New Combinations in Selected Families and Genera. Plant. 2025, 13(2), 76-99. doi: 10.11648/j.plant.20251302.14

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    Feng Z. New Ranks and New Combinations in Selected Families and Genera. Plant. 2025;13(2):76-99. doi: 10.11648/j.plant.20251302.14

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  • @article{10.11648/j.plant.20251302.14,
      author = {Zhen-Hao Feng},
      title = {New Ranks and New Combinations in Selected Families and Genera
    },
      journal = {Plant},
      volume = {13},
      number = {2},
      pages = {76-99},
      doi = {10.11648/j.plant.20251302.14},
      url = {https://doi.org/10.11648/j.plant.20251302.14},
      eprint = {https://article.sciencepublishinggroup.com/pdf/10.11648.j.plant.20251302.14},
      abstract = {Recent advancements in molecular phylogenetics have significantly reshaped infrageneric classifications across vascular plant genera, yet nomenclature within these ranks remains understudied. This article addresses critical gaps in the taxonomy of selected plant families by introducing nomenclatural novelties aligned with modern systematic frameworks. The study is structured into three interrelated sections. First, updated infrageneric classifications are proposed for six genera, i. e., Calanthe, Docynia, Epimedium, Eriolobus, Galanthus, and Nymphaea, integrating morphological and molecular evidence to resolve longstanding taxonomic ambiguities. For instance, formerly segregated genera Cephalantheropsis, Gastrorchis, and Phaius is herein newly treated as subgenera under the broadly circumscribed Calanthe, and Paraphaius is treated as a synonym of Calanthe subgen. Hecabe; while Docynia is retained as distinct from Malus due to its unique ecological and evolutionary adaptations. The second section validates 373 new nothoranks across 58 genera from 29 families of pteridophytes and spermatophytes: Amaranthaceae, Amaryllidaceae, Anemiaceae, Asparagaceae, Asphodelaceae, Asteraceae, Athyriaceae, Berberidaceae, Cistaceae, Crassulaceae, Dryopteridaceae, Heliconiaceae, Hydrangeaceae, Hypericaceae, Iridaceae, Lamiaceae, Lentibulariaceae, Marattiaceae, Musaceae, Nymphaeaceae, Orchidaceae, Papaveraceae, Plantaginaceae, Plumbaginaceae, Ranunculaceae, Rosaceae, Saxifragaceae, Verbenaceae and Violaceae. These include novel nothosections, nothosubsections, nothoseries, and nothosubgenera to accommodate hybrids previously lacking suitable placement. The third section establishes three new nothogenera and 28 new combinations across nine families, namely Apocynaceae, Athyriaceae, Convolvulaceae, Lamiaceae, Orchidaceae, Poaceae, Rosaceae, Saxifragaceae and Thelypteridaceae, resolving taxonomic inconsistencies arising from recent phylogenetic insights. 
    },
     year = {2025}
    }
    

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  • TY  - JOUR
    T1  - New Ranks and New Combinations in Selected Families and Genera
    
    AU  - Zhen-Hao Feng
    Y1  - 2025/06/20
    PY  - 2025
    N1  - https://doi.org/10.11648/j.plant.20251302.14
    DO  - 10.11648/j.plant.20251302.14
    T2  - Plant
    JF  - Plant
    JO  - Plant
    SP  - 76
    EP  - 99
    PB  - Science Publishing Group
    SN  - 2331-0677
    UR  - https://doi.org/10.11648/j.plant.20251302.14
    AB  - Recent advancements in molecular phylogenetics have significantly reshaped infrageneric classifications across vascular plant genera, yet nomenclature within these ranks remains understudied. This article addresses critical gaps in the taxonomy of selected plant families by introducing nomenclatural novelties aligned with modern systematic frameworks. The study is structured into three interrelated sections. First, updated infrageneric classifications are proposed for six genera, i. e., Calanthe, Docynia, Epimedium, Eriolobus, Galanthus, and Nymphaea, integrating morphological and molecular evidence to resolve longstanding taxonomic ambiguities. For instance, formerly segregated genera Cephalantheropsis, Gastrorchis, and Phaius is herein newly treated as subgenera under the broadly circumscribed Calanthe, and Paraphaius is treated as a synonym of Calanthe subgen. Hecabe; while Docynia is retained as distinct from Malus due to its unique ecological and evolutionary adaptations. The second section validates 373 new nothoranks across 58 genera from 29 families of pteridophytes and spermatophytes: Amaranthaceae, Amaryllidaceae, Anemiaceae, Asparagaceae, Asphodelaceae, Asteraceae, Athyriaceae, Berberidaceae, Cistaceae, Crassulaceae, Dryopteridaceae, Heliconiaceae, Hydrangeaceae, Hypericaceae, Iridaceae, Lamiaceae, Lentibulariaceae, Marattiaceae, Musaceae, Nymphaeaceae, Orchidaceae, Papaveraceae, Plantaginaceae, Plumbaginaceae, Ranunculaceae, Rosaceae, Saxifragaceae, Verbenaceae and Violaceae. These include novel nothosections, nothosubsections, nothoseries, and nothosubgenera to accommodate hybrids previously lacking suitable placement. The third section establishes three new nothogenera and 28 new combinations across nine families, namely Apocynaceae, Athyriaceae, Convolvulaceae, Lamiaceae, Orchidaceae, Poaceae, Rosaceae, Saxifragaceae and Thelypteridaceae, resolving taxonomic inconsistencies arising from recent phylogenetic insights. 
    
    VL  - 13
    IS  - 2
    ER  - 

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