3. Nomenclatural Treatment
3.1. Updated Infrageneric Classification for Six Genera
Amaryllidaceae
Galanthus L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[16] | Rønsted, N., Zubov, D., Bruun-Lund, S., Davis, A. P. Snowdrops falling slowly into place: An improved phylogeny for Galanthus (Amaryllidaceae). Molec. Phylogen. Evol. 2013, 69(1), 205-217. https://doi.org/10.1016/j.ympev.2013.05.019 |
[21] | Davis, A. P. The genus Galanthus. Timber Press; 1999, pp. 297. |
[2, 16, 21]
Remark: The morphology-based infrageneric classification of
Galanthus is not fully consistent with phylogenetic findings. Based on phylogenetic results
[16] | Rønsted, N., Zubov, D., Bruun-Lund, S., Davis, A. P. Snowdrops falling slowly into place: An improved phylogeny for Galanthus (Amaryllidaceae). Molec. Phylogen. Evol. 2013, 69(1), 205-217. https://doi.org/10.1016/j.ympev.2013.05.019 |
[16]
,
Galanthus can be divided into seven series, which are corresponding to its seven well-supported clades: ser.
Latifolii Kem.-Nath. (
Platyphyllus clade), ser.
Trojanenses Z. H. Feng (
Trojanus clade), ser.
Ikarienses Z. H. Feng (
Ikariae clade), ser.
Elwesiani Kem.-Nath. ex A. P. Khokhr. (
Elwesii clade), ser.
Galanthus (
Nivalis clade), ser.
Woronowii Kem.-Nath. (
Woronowii clade) and ser.
Alpini Z. H. Feng (
Alpinus clade).
Galanthus ser. Latifolii Kem.-Nath. (1947).
Galanthus ser. Trojanenses Z. H. Feng, ser. nov. Type: Galanthus trojanus A. P. Davis & Özhatay.
Diagnosis: Bulbs ± ovoid or spherical; vernation applanate; leaves linear, sometimes slightly oblanceolate, medium to dark green; inner perianth segments with an apical notch, and a single green mark at the apex, or sometimes with two eye-like marks at the base. Monotypic.
Galanthus ser. Ikarienses Z. H. Feng, ser. nov. Type: Galanthus ikariae Baker.
Diagnosis: Bulbs ± ovoid or spherical; vernation supervoloute; leaves broad-linear to narrowly oblanceolate, light to medium green; inner perianth segments with an apical notch, and a single bold green mark (circa half the size of the segment) at the apex. Monotypic.
Galanthus ser. Elwesiani Kem.-Nath. ex A. P. Khokhr. (1966).
Galanthus ser. Galanthus.
Galanthus ser. Woronowii Kem.-Nath. (1947).
Galanthus ser. Alpini Z. H. Feng, ser. nov. Type: Galanthus alpinus Sosnowsky.
Diagnosis: Bulbs ± ovoid or spherical; vernation supervolute or applanate (only G. angustifolius); leaves linear (G. angustifolius, G. koenenianus) to narrowly oblanceolate, glaucous or greenglaucescent (G. transcaucasicus); inner perianth segments with an apical notch, and a single green mark at the apex, sometimes with a faint yellowish or green mark at the base (only G. koenenianus). Four species: G. alpinus, G. angustifolius, G. koenenianus and G. transcaucasicus.
Berberidaceae
Epimedium L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[13] | He, S.-Z. The genus Epimedium of China in colour. Guizhou Science and Technology Publishing House, Guizhou, China; 2014, pp. 203. |
[14] | Zhang, Y., Yang, L., Chen, J., Sun, W., Wang, Y. Taxonomic and phylogenetic analysis of Epimedium L. based on amplified fragment length polymorphisms. Sci. Hort. (Amsterdam) 2014, 170, 284-292. https://doi.org/10.1016/j.scienta.2014.02.025 |
[2, 13, 14]
Remark: The serial classification of
Epimedium sect.
Diphyllon is controversial by different authors using floral spur, number of cauline leaf, rhizome or geographical distribution.
[13] | He, S.-Z. The genus Epimedium of China in colour. Guizhou Science and Technology Publishing House, Guizhou, China; 2014, pp. 203. |
[13]
According to the subsequent molecular findings, Chinese sect.
Diphyllon was divided into five well-supported clades related to flower morphology except that five species were either isolated or formed a general polytomy.
[14] | Zhang, Y., Yang, L., Chen, J., Sun, W., Wang, Y. Taxonomic and phylogenetic analysis of Epimedium L. based on amplified fragment length polymorphisms. Sci. Hort. (Amsterdam) 2014, 170, 284-292. https://doi.org/10.1016/j.scienta.2014.02.025 |
[14]
Epimedium subgen. Rhizophyllum (Fisch. & C. A. Mey.) Stearn (2002).
Epimedium subgen. Epimedium
Epimedium sect. Diphyllon (Kom.) Stearn (2002).
Epimedium ser. Calcar (S. Z. He) Z. H. Feng, stat. nov. — Basionym: Epimedium sect. Calcar S. Z. He, Genus Epimedium China: 28. 2014.
Epimedium ser. Davidianae Stearn (2002).
Epimedium ser. Diphyllon Kom. (1908).
Epimedium ser. Dolichocera Stearn (1938).
Epimedium ser. Fargesii Z. H. Feng, ser. nov. Type: Epimedium fargesii Franch.
Diagnosis: Flowers small but slightly larger than those of Epimedium ser. Diphyllon, stamens conspicuously protruded, 8-10 mm long. Four species: E. dewuense, E. dolichostemon, E. fargesii and E. qingchengshanense.
Epimedium ser. Leptorrhizi Z. H. Feng, ser. nov. Type: Epimedium leptorrhizum Stearn.
Diagnosis: Rhizome long-creeping, 1-2 mm thick, with internodes sometimes as much as 20 cm long. Petioles furnished with spreading reddish hairs; lateral leaflet often pruinose abaxially. Monotypic.
Epimedium ser. Stellulati Z. H. Feng, ser. nov. Type: Epimedium stellulatum Stearn.
Diagnosis: Inflorescence compound, loose, circa 15-20 cm long, pedicels with numerous glandular hairs. Flowers small, Petals much shorter than inner sepals, with blunt nectariferous spur. Monotypic.
Epimedium sect. Epimedium
Epimedium sect. Macroceras C. Morren & Decne. (1834).
Epimedium sect. Polyphyllon (Kom.) Stearn (2002).
Nymphaeaceae
Nymphaea L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[17] | Conard, H. S. The waterlilies: A monograph of the genus Nymphaea. Carnegie Institution of Washington; 1905, pp. 279. https://doi.org/10.5962/bhl.title.23557 |
[18] | Löhne, C., Yoo, M.-J., Borsch, T., Wiersema, J., Wilde, V., Bell, C. D., Barthlott, W., Soltis, D. E., Soltis, P. S. Biogeography of Nymphaeales: Extant patterns and historical events. Taxon 2008, 57(4), 1123-1146. https://doi.org/10.1002/tax.574008 |
[2, 17, 18]
Remark: Tradionally,
Nymphaea is divided into five subgenera based on morphological traits, such as filament appearance, sterile appendage and carpel walls free or fused.
Jacobs, in 2007, added a new subgenus
Confluentes, characterized by flowers with a distinct gap between petals and stamens, leaf margins toothed and seeds rather large. According to the subsequent molecular findings
[18] | Löhne, C., Yoo, M.-J., Borsch, T., Wiersema, J., Wilde, V., Bell, C. D., Barthlott, W., Soltis, D. E., Soltis, P. S. Biogeography of Nymphaeales: Extant patterns and historical events. Taxon 2008, 57(4), 1123-1146. https://doi.org/10.1002/tax.574008 |
[18]
, it is warrant to reduce
Confluentes as a section within subgen.
Anecphya s. lat.
Nymphaea subgen. Anecphya (Casp.) Conard (1905).
Nymphaea sect. Confluentes (S. W. L. Jacobs) Bing Liu ex Zi Jun Li & Z. H. Feng, stat. nov. — Basionym: Nymphaea subgen. Confluentes S. W. L. Jacobs, Fl. Australia 2: 458. 2007.
Nymphaea sect. Anecphya (Casp.) Bing Liu ex Zi Jun Li & Z. H. Feng, stat. nov. — Basionym: Nymphaea subsect. Anecphya Casp., Ann. Mus. Bot. Lugduno-Batavi 2: 247. 1866.
Nymphaea subgen. Brachyceras (Casp.) Conard (1905).
Nymphaea subgen. Nymphaea
Nymphaea sect. Xanthantha (Casp.) Wiersema (1996).
Nymphaea sect. Nymphaea
Nymphaea subgen. Lotos (DC.) Conard (1905).
Nymphaea subgen. Hydrocallis (Planch.) Conard (1905).
Orchidaceae
Calanthe R. Br.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[6] | Clayton, D., Cribb, P. The genus Calanthe. Natural History Publications (Borneo) in association with Royal Botanic Gardens, Kew, Kota Kinabalu; 2013, pp. 413. |
[7] | Zhai, J.-W., Zhang, G.-Q., Li, L., Wang, M., Chen, L.-J., Chung, S.-W., Rodríguez, F. J., Francisco-Ortega, J., Lan, S.-R., Xing, F.-W., Liu, Z.-J. A new phylogenetic analysis sheds new light on the relationships in the Calanthe alliance (Orchidaceae) in China. Molec. Phylogen. Evol. 2014, 77, 216-222. https://doi.org/10.1016/j.ympev.2014.04.005 |
[8] | Chase, M. W., Christenhusz, M. J. M., Schuiteman, A. Expansion of Calanthe to include the species of Cephalantheropsis, Gastrorchis and Phaius (Collabieae; Orchidaceae). Phytotaxa 2020, 472(2), 159-168. https://doi.org/10.11646/phytotaxa.472.2.6 |
[9] | Chase, M. W., Christenhusz, M. J. M., Schuiteman, A. (2782) Proposal to conserve Calanthe, nom. cons., against the additional names Phaius, Cyanorkis, and Gastorkis (Orchidaceae, Collabieae). Taxon 2020, 69(6), 1364-1365. https://doi.org/10.1002/tax.12396 |
[10] | Wilson, K. L. Report of the General Committee: 26. Taxon 2023, 72(4), 905-907. https://doi.org/10.1002/tax.12994 |
[11] | Applequist, W. L. Report of the Nomenclature Committee for Vascular Plants: 73. Taxon 2023, 72(1), 179-204. https://doi.org/10.1002/tax.12871 |
[2, 6-11]
Remark: In
Calanthe, seven major subclades are found by molecular phylogenetic analyses covering the major infrageneric groups in
Calanthe and
Phaius plus representative species of allied genera. Among which, subclade (6) was elevated to generic level as
Paraphaius.
[6] | Clayton, D., Cribb, P. The genus Calanthe. Natural History Publications (Borneo) in association with Royal Botanic Gardens, Kew, Kota Kinabalu; 2013, pp. 413. |
[7] | Zhai, J.-W., Zhang, G.-Q., Li, L., Wang, M., Chen, L.-J., Chung, S.-W., Rodríguez, F. J., Francisco-Ortega, J., Lan, S.-R., Xing, F.-W., Liu, Z.-J. A new phylogenetic analysis sheds new light on the relationships in the Calanthe alliance (Orchidaceae) in China. Molec. Phylogen. Evol. 2014, 77, 216-222. https://doi.org/10.1016/j.ympev.2014.04.005 |
[6, 7]
However, Chase et al.
[8] | Chase, M. W., Christenhusz, M. J. M., Schuiteman, A. Expansion of Calanthe to include the species of Cephalantheropsis, Gastrorchis and Phaius (Collabieae; Orchidaceae). Phytotaxa 2020, 472(2), 159-168. https://doi.org/10.11646/phytotaxa.472.2.6 |
[8]
sought a broader delimitation of
Calanthe to include the species of
Cephalantheropsis,
Gastrorchis and
Phaius because “segregate new genera from both
Phaius and
Calanthe … results in generic concepts for which there are no reasonable morphological differences”, and since
Calanthe is conserved against
Phaius [9] | Chase, M. W., Christenhusz, M. J. M., Schuiteman, A. (2782) Proposal to conserve Calanthe, nom. cons., against the additional names Phaius, Cyanorkis, and Gastorkis (Orchidaceae, Collabieae). Taxon 2020, 69(6), 1364-1365. https://doi.org/10.1002/tax.12396 |
[10] | Wilson, K. L. Report of the General Committee: 26. Taxon 2023, 72(4), 905-907. https://doi.org/10.1002/tax.12994 |
[11] | Applequist, W. L. Report of the Nomenclature Committee for Vascular Plants: 73. Taxon 2023, 72(1), 179-204. https://doi.org/10.1002/tax.12871 |
[9-11]
, a broad sense of
Calanthe is here adopted with an updated infrageneric classification.
Calanthe subgen. Calanthe
Calanthe sect. Aceratochilus Schltr. (1912).
Calanthe sect. Alpinocalanthe J. W. Zhai, Z. J. Liu & F. W. Xing (2014).
Calanthe sect. Ghiesbreghtia (A. Rich. & Galeotti) Schltr. (1912).
Calanthe sect. Monophylla D. A. Clayton & P. J. Cribb (2013).
Calanthe sect. Puberula J. W. Zhai, Z. J. Liu & F. W. Xing (2014).
Calanthe sect. Tricarinata J. W. Zhai, Z. J. Liu & F. W. Xing (2014).
Calanthe sect. Verna Kakadzu (1994).
Calanthe subgen. Cephalantheropsis (Guillaumin) Z. H. Feng, comb. & stat. nov. — Basionym: Cephalantheropsis Guillaumin, Bull. Mus. Natl. Hist. Nat., sér. 2, 32: 188. 1960.
Calanthe subgen. Gastrorchis (Schltr.) Z. H. Feng, comb. & stat. nov. — Basionym: Gastrorchis Schltr., Repert. Spec. Nov. Regni Veg. Beih. 33: 167. 1924.
Calanthe subgen. Hecabe (Raf.) Z. H. Feng, comb. & stat. nov. — Basionym: Hecabe Raf., Fl. Tellur. 4: 44. 1838. — Synonym: Paraphaius J. W. Zhai & F. W. Xing, Molec. Phylogen. Evol. 77: 221. 2014.
Calanthe subgen. Phaius (Lour.) Z. H. Feng, comb. & stat. nov. — Basionym: Phaius Lour., Fl. Cochinch. 2: 517, 529. 1790.
Calanthe subgen. Preptanthe (Rchb. f.) Schltr. (1912).
Calanthe subgen. Styloglossum Kakadzu (1994).
Rosaceae
Docynia Decne.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[12] | Liu, G.-N., Ma, D.-K., Xu, C., Huang, J., Ge, B.-J., Luo, Q., Wei, Y., Liu, B.-B. Malus includes Docynia (Maleae, Rosaceae): Evidence from phylogenomics and morphology. PhytoKeys 2023, 229, 47-60. https://doi.org/10.3897/phytokeys.229.103888 |
[2, 12]
Remark: The
Docynia-
Docyniopsis clade has a conflicting phylogenetic position in the nuclear and plastid trees, indicating that the evolutionary relationships are not clear-cut. Liu et al.
[12] | Liu, G.-N., Ma, D.-K., Xu, C., Huang, J., Ge, B.-J., Luo, Q., Wei, Y., Liu, B.-B. Malus includes Docynia (Maleae, Rosaceae): Evidence from phylogenomics and morphology. PhytoKeys 2023, 229, 47-60. https://doi.org/10.3897/phytokeys.229.103888 |
[12]
argued that “narrow generic concepts may be impractical for use”, however, this does not necessarily justify lumping because taxonomic circumscription should consider the broader implications for conservation, ecology, and evolution, simply lumping
Docynia into
Malus based on obviously unconvincing evidence may benifit the nomenclature but disrupt the understanding of the unique evolutionary adaptations and ecological roles of
Docynia.
Docynia is here kept as an independent genus, and is divided into two sections based on morpho-phylogenetic evidences: sect.
Docynia encompasses
D. delavayi,
D. indica and
D. longiunguis, and sect.
Paradocynia contains only
D. doumeri.
Docynia sect. Docynia
Docynia sect. Paradocynia Su Liu ex Z. H. Feng, sect. nov. Type: Docynia doumeri (Bois) C. K. Schneid.
Diagnosis: Lamina oblong to ovate, base rounded or cuneate, apex acute or acuminate, pubescent when young then glabrescent, or glabrous; styles 5, pubescent at base; pome ca 2.5‒5.5 cm in diameter, subglobose, yellow to red. Monotypic.
Eriolobus (DC.) M. Roem.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[12] | Liu, G.-N., Ma, D.-K., Xu, C., Huang, J., Ge, B.-J., Luo, Q., Wei, Y., Liu, B.-B. Malus includes Docynia (Maleae, Rosaceae): Evidence from phylogenomics and morphology. PhytoKeys 2023, 229, 47-60. https://doi.org/10.3897/phytokeys.229.103888 |
[15] | Wang, H., Li, X.-Y., Jiang, Y., Jin, Z.-T., Ma, D.-K., Liu, B., Xu, C., Ge, B.-J., Wang, T., Fan, Q., Jin, S.-H., Liu, G.-N., Liu, B.-B. Refining the phylogeny and taxonomy of the apple tribe Maleae (Rosaceae): Insights from phylogenomic analyses of 563 plastomes and a taxonomic synopsis of Photinia and its allies in the Old World. PhytoKeys 2024, 242, 161-227. https://doi.org/10.3897/phytokeys.242.117481 |
[2, 12, 15]
Remark: Although Liu et al.
[12] | Liu, G.-N., Ma, D.-K., Xu, C., Huang, J., Ge, B.-J., Luo, Q., Wei, Y., Liu, B.-B. Malus includes Docynia (Maleae, Rosaceae): Evidence from phylogenomics and morphology. PhytoKeys 2023, 229, 47-60. https://doi.org/10.3897/phytokeys.229.103888 |
[12]
argued that the transferring of
Docynia to the genus
Eriolobus is untenable, they, unfortunately, did not take the similar choice as another line of evidence to keep
Eriolobus apart from
Malus s. s. According to Wang et al.
[15] | Wang, H., Li, X.-Y., Jiang, Y., Jin, Z.-T., Ma, D.-K., Liu, B., Xu, C., Ge, B.-J., Wang, T., Fan, Q., Jin, S.-H., Liu, G.-N., Liu, B.-B. Refining the phylogeny and taxonomy of the apple tribe Maleae (Rosaceae): Insights from phylogenomic analyses of 563 plastomes and a taxonomic synopsis of Photinia and its allies in the Old World. PhytoKeys 2024, 242, 161-227. https://doi.org/10.3897/phytokeys.242.117481 |
[15]
and their previous research,
Malus is dyphyletic, the North American crab apples clade being distant to the type clade, proving that it is an irrational choice to oversimply lumping
Eriolobus into
Malus.
Eriolobus is here recognized as independent genus and is divided into three sections based on morpho-phylogenetic evidences, sect.
Eriolobus includes only the type species
E. trilobatus, sect.
Chloromeles contains
E. angustifolius,
E. coronarius and
E. ioensis, while sect.
Florentinae solely embraces
E. florentinus.
Eriolobus sect. Chloromeles (Decne.) Su Liu ex Z. H. Feng, comb. & stat. nov. — Basionym: Malus subgen. Chloromeles Decne., Nouv. Arch. Mus. Hist. Nat. Paris 10: 155. 1874. Type: Eriolobus angustifolius (Aiton) Su Liu ex Z. H. Feng.
Eriolobus sect. Eriolobus
Eriolobus sect. Florentinae (Rehder) Su Liu ex Z. H. Feng, comb. & stat. nov. — Basionym: Malus subsect. Florentinae Rehder, J. Arnold Arbor. 2: 48. 1920. Type: Eriolobus florentinus (Zuccagni) Stapf.
3.2. New Nothoranks of Pteridophytes and Spermatophytes
Amaranthaceae
Amaranthus nothosect. Blorion Z. H. Feng, nothosect. nov. = Amaranthus sect. Blitopsis Dumort. × Amaranthus sect. Pentamorion (Beck) Mosyakin & K. R. Robertson.
Amaranthus nothosect. Pentidium Z. H. Feng, nothosect. nov. = Amaranthus sect. Pentamorion (Beck) Mosyakin & K. R. Robertson × Amaranthus sect. Pyxidium Moq.
Amaranthus nothosect. Pyxopsis Z. H. Feng, nothosect. nov. = Amaranthus sect. Blitopsis Dumort. × Amaranthus sect. Pyxidium Moq.
Amaranthus nothosubgen. Amarida Z. H. Feng, nothosubgen. nov. = Amaranthus subgen. Amaranthus × Amaranthus subgen. Acnida (L.) Aellen ex K. R. Robertson.
Amaranthus nothosubsect. Hybramaranthus Z. H. Feng, nothosubsect. nov. = Amaranthus subsect. Amaranthus × Amaranthus subsect. Hybrida Mosyakin & K. R. Robertson.
Amaranthus nothosubsect. Hynida Z. H. Feng, nothosubsect. nov. = Amaranthus sect. Acnida (L.) Mosyakin & K. R. Robertson × Amaranthus subsect. Hybrida Mosyakin & K. R. Robertson.
Amaryllidaceae
Galanthus L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[16] | Rønsted, N., Zubov, D., Bruun-Lund, S., Davis, A. P. Snowdrops falling slowly into place: An improved phylogeny for Galanthus (Amaryllidaceae). Molec. Phylogen. Evol. 2013, 69(1), 205-217. https://doi.org/10.1016/j.ympev.2013.05.019 |
[21] | Davis, A. P. The genus Galanthus. Timber Press; 1999, pp. 297. |
[2, 16, 21]
Galanthus nothoser. Woropini Z. H. Feng, nothoser. nov. = Galanthus ser. Alpini Z. H. Feng × Galanthus ser. Woronowii Kem.-Nath.
Anemiaceae
Anemia nothosect. Anemutae Z. H. Feng, nothosect. nov. = Anemia sect. Anemia × Anemia sect. Hirsutae Prantl.
Anemia nothosect. Aninae Z. H. Feng, nothosect. nov. = Anemia sect. Anemia × Anemia sect. Collinae Prantl.
Anemia nothosect. Cosutae Z. H. Feng, nothosect. nov. = Anemia sect. Collinae Prantl. × Anemia sect. Hirsutae Prantl.
Anemia nothosect. Tomemia Z. H. Feng, nothosect. nov. = Anemia sect. Anemia × Anemia sect. Tomentosae Prantl.
Asparagaceae
Agave L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[23] | Thiede, J., Smith, G. F., Eggli, U. Infrageneric classification of Agave L. (Asparagaceae: Agavoideae / Agavaceae): A nomenclatural assessment and updated classification at the rank of section, with new combinations. Bradleya 2019, 37, 240-264. https://doi.org/10.25223/brad.n37.2019.a22 |
[2, 23]
Agave nothosect. Deslorae Z. H. Feng, nothosect. nov. = Agave sect. Deserticolae (Trel.) R. H. Webb & G. D. Starr × Agave sect. Parviflorae Hochstätter.
Agave nothosect. Hetemoratae Z. H. Feng, nothosect. nov. = Agave sect. Heteracanthae Salm-Dyck × Agave sect. Marmoratae (A. Berger) Thiede & Gideon F. Sm.
Agave nothosect. Heticracanthae Z. H. Feng, nothosect. nov. = Agave sect. Heteracanthae Salm-Dyck × Agave sect. Micracanthae Salm-Dyck.
Agave nothosect. Hibacanthae Z. H. Feng, nothosect. nov. = Agave sect. Heteracanthae Salm-Dyck × Agave sect. Hibernicae Hochstätter.
Agave nothosect. Littanthae Z. H. Feng, nothosect. nov. = Agave sect. Littaea (Tagl.) Benth. × Agave sect. Micracanthae Salm-Dyck.
Agave nothosect. Micragave Z. H. Feng, nothosect. nov. = Agave sect. Agave × Agave sect. Micracanthae Salm-Dyck.
Agave nothosect. Parranthae Z. H. Feng, nothosect. nov. = Agave sect. Heteracanthae Salm-Dyck × Agave sect. Parryanae Hochstätter.
Agave nothosect. Parvipalae Z. H. Feng, nothosect. nov. = Agave sect. Ditepalae Hochstätter × Agave sect. Parviflorae Hochstätter.
Agave nothoser. Bravanthes (Cif. & Giacom.) Z. H. Feng, comb. & stat. nov. = Agave ser. Bravoa (Lex.) Thiede & Gideon F. Sm. × Agave ser. Polianthes (L.) Thiede & Gideon F. Sm. — Basionym: × Bravanthes Cif. & Giacom., Nomencl. Fl. Ital. Pt. 1: 134. 1950.
Agave nothosubgen. Agavaea Z. H. Feng, nothosubgen. nov. = Agave subgen. Agave × Agave subgen. Littaea (Tagl.) Baker.
Asphodelaceae
Hemerocallis L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[24] | Murakami, S., Takayama, K., Fuse, S., Hirota, S. K., Koi, S., Ideno, T., Yamamoto, T., Tamura, M. N. Recircumscription of sections of Hemerocallis (Asphodelaceae) from Japan and adjacent regions based on MIG-seq Data. Acta Phytotax. Geobot. 2020, 71(1), 1-11. https://doi.org/10.18942/apg.201913 |
[2, 24]
Hemerocallis nothosect. Hemerulvae Z. H. Feng & H. N. Sun, nothosect. nov. = Hemerocallis sect. Fulvae Nakai × Hemerocallis sect. Hemerocallis.
Asteraceae
Arctium L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[25] | López-Vinyallonga, S., Romaschenko, K., Susanna, A., Garcia-Jacas, N. Systematics of the Arctioid group: Disentangling Arctium and Cousinia (Cardueae, Carduinae). Taxon 2011, 60(2), 539-554. https://doi.org/10.1002/tax.602020 |
[2, 25]
Arctium nothosect. Pecarctium Z. H. Feng, nothosect. nov. = Arctium sect. Pectinata (C. Winkl.) S. López, Romasch., Susanna & N. Garcia × Arctium sect. Pseudarctium (Juz.) Duist.
Artemisia L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[26] | Jiao, B.-H., Chen, C., Wei, M., Niu, G.-H., Zheng, J.-Y., Zhang, G.-J., Shen, J.-H., Vitales, D., Vallès, J., Verloove, F., Erst, A. S., Soejima, A., Mehregan, I., Kokubugata, G., Chung, G.-Y., Ge, X.-J., Gao, L.-M., Yuan, Y., Joly, C., Jabbour, F., Wang, W., Shultz, L. M., Gao, T.-G. Phylogenomics and morphological evolution of the mega-diverse genus Artemisia (Asteraceae: Anthemideae): implications for its circumscription and infrageneric taxonomy. Ann. Bot. (Oxford) 2023, 131(5), 867-883. https://doi.org/10.1093/aob/mcad051 |
[2, 26]
Artemisia nothosubgen. Artinthium Z. H. Feng, nothosubgen. nov. = Artemisia subgen. Absinthium (Mill.) Less. × Artemisia subgen. Artemisia.
Artemisia nothosubgen. Dracunthium Z. H. Feng, nothosubgen. nov. = Artemisia subgen. Absinthium (Mill.) Less. × Artemisia subgen. Dracunculus (Besser) Rydb.
Artemisia nothosubgen. Ponculus Z. H. Feng, nothosubgen. nov. = Artemisia subgen. Dracunculus (Besser) Rydb. × Artemisia subgen. Ponticae B. H. Jiao ex T. G. Gao.
Artemisia nothosubgen. Sesinthium Z. H. Feng, nothosubgen. nov. = Artemisia subgen. Absinthium (Mill.) Less. × Artemisia subgen. Seriphidium Besser ex Less.
Baccharis L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[27] | Heiden, G., Antonelli, A., Pirani, J. R. A novel phylogenetic infrageneric classification of Baccharis (Asteraceae: Astereae), a highly diversified American genus. Taxon 2019, 68(5), 1048-1081. https://doi.org/10.1002/tax.12128 |
[2, 27]
Baccharis nothosect. Pedifoliae Z. H. Feng, nothosect. nov. = Baccharis sect. Cuneifoliae DC. × Baccharis sect. Pedicellatae Heering.
Cousinia Cass.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[28] | Tscherneva, O. New supraspecific taxa of the genus Cousinia (Asteraceae). Bot. Zhurn. (Moscow & Leningrad) 1988, 73(4), 594-597. |
[29] | Mehregan, I., Kadereit, J. W. Taxonomic revision of Cousinia sect. Cynaroideae (Asteraceae, Cardueae). Willdenowia 2008, 38(2), 293-362. https://doi.org/10.3372/wi.38.38201 |
[30] | Mehregan, I., Kadereit, J. W. The role of hybridization in the evolution of Cousinia s. str. (Asteraceae, Cardueae). Willdenowia 2009, 39(1), 35-47. https://doi.org/10.3372/wi.39.39102 |
[31] | Sennikov, A. N. A revision of Cousinia sections Alpinae (syn. Carduncellus), Subappendiculatae and Tianschanicae (Asteraceae) in the Kirghizian Tian-Shan and the neighbouring territories. Phytotaxa 2013, 5(1), 1-30. https://doi.org/10.11646/phytotaxa. 5.1.1 |
[2, 28-31]
Cousinia nothosect. Alpiopsis Z. H. Feng, nothosect. nov. = Cousinia sect. Alpinae Bunge × Cousinia sect. Jurineopsis (Juz.) Tscherneva.
Cousinia nothosect. Coronopsis Z. H. Feng, nothosect. nov. = Cousinia sect. Coronophora (Juz.) Rech. f. × Cousinia sect. Jurineopsis (Juz.) Tscherneva.
Cousinia nothosect. Cynarocaules Z. H. Feng, nothosect. nov. = Cousinia sect. Cynaroideae Bunge × Cousinia sect. Leiocaules Bunge.
Cousinia nothosect. Cynaroferae Z. H. Feng, nothosect. nov. = Cousinia sect. Cynaroideae Bunge × Cousinia sect. Pugioniferae Bunge.
Cousinia nothosect. Pulchecarpae Z. H. Feng, nothosect. nov. = Cousinia sect. Microcarpae Bunge × Cousinia sect. Pulchellae Rech. f.
Cousinia nothosect. Pulchepinae Z. H. Feng, nothosect. nov. = Cousinia sect. Alpinae Bunge × Cousinia sect. Pulchellae Rech. f.
Cousinia nothosect. Serracousinia Z. H. Feng, nothosect. nov. = Cousinia sect. Eriocousinia Tscherneva × Cousinia sect. Serratuloides Bunge.
Cousinia nothosect. Sphaeroides Z. H. Feng, nothosect. nov. = Cousinia sect. Serratuloides Bunge × Cousinia sect. Sphaerocephalae Bunge.
Liatris Gaertn. ex Schreb.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[32] | Nesom, G. L. Infrageneric classification of Liatris (Asteraceae: Eupatorieae). Sida 2005, 21, 1305-1321. |
[2, 32]
Liatris nothosect. Gramago Z. H. Feng, nothosect. nov. = Liatris sect. Graminifolium G. L. Nesom × Liatris sect. Suprago (Gaertn.) DC.
Liatris nothosect. Liafolium Z. H. Feng, nothosect. nov. = Liatris sect. Graminifolium G. L. Nesom × Liatris sect. Liatris.
Liatris nothosect. Liprago Z. H. Feng, nothosect. nov. = Liatris sect. Liatris × Liatris sect. Suprago G. L. Nesom.
Liatris nothosect. Lirago Z. H. Feng, nothosect. nov. = Liatris sect. Liatris × Liatris sect. Vorago G. L. Nesom.
Liatris nothoser. Elerago Z. H. Feng, nothoser. nov. = Liatris ser. Elegantes (Alexander) Gaiser ex G. L. Nesom × Liatris sect. Vorago G. L. Nesom.
Liatris nothoser. Liprago (Z. H. Feng) Z. H. Feng, stat. nov. = Liatris ser. Liatris × Liatris sect. Suprago G. L. Nesom. — Basionym: Liatris nothosect. Liprago Z. H. Feng, this publication.
Liatris nothoser. Lirago (Z. H. Feng) Z. H. Feng, stat. nov. = Liatris ser. Liatris × Liatris sect. Vorago G. L. Nesom. — Basionym: Liatris nothosect. Lirago Z. H. Feng, this publication.
Liatris nothoser. Punosae Z. H. Feng, nothoser. nov. = Liatris ser. Punctatae (Alexander) Gaiser ex Fernald × Liatris ser. Scariosae (Alexander) Gaiser ex Fernald.
Liatris nothoser. Scrago Z. H. Feng, nothoser. nov. = Liatris ser. Scariosae (Alexander) Gaiser ex Fernald × Liatris sect. Suprago (Gaertn.) DC.
Athyriaceae
Athyrium Roth
[33] | Wei, R., Ebihara, A., Zhu, Y.-M., Zhao, C.-F., Hennequin, S., Zhang, X.-C. A total-evidence phylogeny of the lady fern genus Athyrium Roth (Athyriaceae) with a new infrageneric classification. Molec. Phylogen. Evol. 2018, 119, 25-36. https://doi.org/10.1016/j.ympev.2017.10.019 |
[33]
Athyrium nothosect. Bistichoides Z. H. Feng, nothosect. nov. = Athyrium sect. Biserrulata (Ching & Y. T. Hsieh) R. Wei & X. C. Zhang × Athyrium sect. Polystichoides Ching & Y. T. Hsieh.
Athyrium nothosect. Otoniana Z. H. Feng, nothosect. nov. = Athyrium sect. Mackinnoniana (Ching & Y. T. Hsieh) Z. R. Wang × Athyrium sect. Otophora Z. R. Wang.
Athyrium nothosect. Otoscentia Z. H. Feng, nothosect. nov. = Athyrium sect. Otophora Z. R. Wang × Athyrium sect. Yokoscentia (Z. R. Wang) R. Wei & X. C. Zhang.
Athyrium nothosect. Polyphora Z. H. Feng, nothosect. nov. = Athyrium sect. Otophora Z. R. Wang × Athyrium sect. Polystichoides Ching & Y. T. Hsieh.
Athyrium nothosect. Yokoniana Z. H. Feng, nothosect. nov. = Athyrium sect. Mackinnoniana (Ching & Y. T. Hsieh) Z. R. Wang × Athyrium sect. Yokoscentia (Z. R. Wang) R. Wei & X. C. Zhang.
Deparia Medik.
[34] | Kuo, L.-Y., Ebihara, A., Hsu, T.-C., Rouhan, G., Huang, Y.-M., Wang, C.-N., Chiou, W.-L., & Kato, M. Infrageneric Revision of the Fern Genus Deparia (Athyriaceae, Aspleniineae, Polypodiales). Syst. Bot. 2018, 43(3), 645-655. https://doi.org/10.1600/036364418X697364 |
[34]
Deparia nothosect. Lunoathyrium Z. H. Feng, nothosect. nov. = Deparia sect. Dryoathyrium (Ching) M. Kato × Deparia sect. Lunathyrium (Koidz.) M. Kato.
Deparia nothosubsect. Depopsis Z. H. Feng, nothosubsect. nov. = Deparia subsect. Athyriopsis (Ching) L. Y. Kuo, M. Kato & W. L. Chiou × Deparia subsect. Deparia.
Deparia nothosubsect. Lunathyriopsis Z. H. Feng, nothosubsect. nov. = Deparia subsect. Athyriopsis (Ching) L. Y. Kuo, M. Kato & W. L. Chiou × Deparia sect. Lunathyrium (Koidz.) M. Kato.
Diplazium Sw.
[35] | Wei, R., Sundue, M., Yang, J., Zhang, X.-C. A sectional classification of Diplazium Sw. (Athyriaceae), based on plastid genomic, nuclear ribosomal, and morphological evidence. Taxon 2025, 74(1), 13-38. https://doi.org/10.1002/tax.13281 |
[35]
Diplazium nothosect. Anisostegia Z. H. Feng, nothosect. nov. = Diplazium sect. Anisogonium (C. Presl) Alderw. × Diplazium sect. Dolichostegia (W. M. Chu & Z. R. He) R. Wei.
Diplazium nothosect. Anisotodia Z. H. Feng, nothosect. nov. = Diplazium sect. Allantodia (R. Br.) R. Wei × Diplazium sect. Anisogonium (C. Presl) Alderw.
Diplazium nothosect. Dolichomatum Z. H. Feng, nothosect. nov. = Diplazium sect. Dolichostegia (W. M. Chu & Z. R. He) R. Wei × Diplazium sect. Longirhizomatum (W. M. Chu & Z. R. He) R. Wei.
Diplazium nothosect. Sibistegia Z. H. Feng, nothosect. nov. = Diplazium subgen. Sibirica R. Wei & X. C. Zhang × Diplazium sect. Dolichostegia (W. M. Chu & Z. R. He) R. Wei.
Diplazium nothosubgen. Sibipteris Z. H. Feng, nothosubgen. nov. = Diplazium subgen. Callipteris (Bory) R. Wei & X. C. Zhang × Diplazium subgen. Sibirica R. Wei & X. C. Zhang.
Berberidaceae
Epimedium L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[13] | He, S.-Z. The genus Epimedium of China in colour. Guizhou Science and Technology Publishing House, Guizhou, China; 2014, pp. 203. |
[14] | Zhang, Y., Yang, L., Chen, J., Sun, W., Wang, Y. Taxonomic and phylogenetic analysis of Epimedium L. based on amplified fragment length polymorphisms. Sci. Hort. (Amsterdam) 2014, 170, 284-292. https://doi.org/10.1016/j.scienta.2014.02.025 |
[2, 13, 14]
Epimedium nothosect. Rhizepimedium (Z. H. Feng) Z. H. Feng, stat. nov. = Epimedium sect. Epimedium × Epimedium subgen. Rhizophyllum (Fisch. & C. A. Mey.) Stearn. — Basionym: Epimedium nothosubgen. Rhizepimedium Z. H. Feng, this publication.
Epimedium nothosect. Rhizoceras Z. H. Feng, nothosect. nov. = Epimedium sect. Macroceras C. Morren & Decne. × Epimedium subgen. Rhizophyllum (Fisch. & C. A. Mey.) Stearn.
Epimedium nothosect. Macrepimedium Z. H. Feng, nothosect. nov. = Epimedium sect. Epimedium × Epimedium sect. Macroceras C. Morren & Decne.
Epimedium nothoser. Dolidianae Z. H. Feng, nothoser. nov. = Epimedium ser. Dolichocera Stearn × Epimedium ser. Davidianae Stearn.
Epimedium nothosubgen. Rhizepimedium Z. H. Feng, nothosubgen. nov. = Epimedium subgen. Epimedium × Epimedium subgen. Rhizophyllum (Fisch. & C. A. Mey.) Stearn.
Cistaceae
Helianthemum Mill.
[36] | Martín-Hernanz, S., Velayos, M., Albaladejo, R. G., Aparicio, A. Systematic implications from a robust phylogenetic reconstruction of the genus Helianthemum (Cistaceae) based on genotyping-by-sequencing (GBS) data. Anales Jard. Bot. Madrid 2021, 78(2), e113. https://doi.org/10.3989/ajbm.2601 |
[36]
Helianthemum nothosect. Lavanthemum Z. H. Feng, nothosect. nov. = Helianthemum sect. Helianthemum × Helianthemum sect. Lavandulaceum G. López.
Helianthemum nothosect. Pseudanthemum Z. H. Feng, nothosect. nov. = Helianthemum sect. Helianthemum × Helianthemum sect. Pseudocistus Dunal.
Helianthemum nothosubgen. Erionthemum Z. H. Feng, nothosubgen. nov. = Helianthemum subgen. Eriocarpum (Dunal) Martín-Hernanz, Velayos, Albaladejo & Aparicio × Helianthemum subgen. Helianthemum.
Helianthemum nothosubgen. Plecanthemum Z. H. Feng, nothosubgen. nov. = Helianthemum subgen. Helianthemum × Helianthemum subgen. Plectolobum Willk.
Crassulaceae
Aichryson Webb & Berthel.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[37] | Bramwell, D. Notes on the taxonomy and nomenclature of the genus Aichryson. Bol. Inst. Nac. Invest. Agron. 1968, 28, 203-213. |
[2, 37]
Aichryson nothosect. Aichrobia Z. H. Feng, nothosect. nov. = Aichryson sect. Aichryson × Aichryson sect. Macrobia Webb & Berthel.
Kalanchoe Adans.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[38] | Smith, G. F. Nomenclature of the partial infrageneric classification proposed for Kalanchoe (Crassulaceae subfam. Kalanchooideae) by René C. J. E. Maire in 1976. Phytotaxa 2020, 468(2), 231-235. https://doi.org/10.11646/phytotaxa.468.2.8 |
[39] | Shtein, R., Smith, G. F. A revision of the climbing kalanchoes (Crassulaceae subfam. Kalanchooideae) of Madagascar including the description of Kalanchoe sect. Invasores and K. ser. Vilana. Phytotaxa 2021, 482(2), 93-120. https://doi.org/10.11646/phytotaxa.482.2.1 |
[40] | Smith, G. F. Kalanchoe sect. Raveta (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1916): A long-overlooked name in K. subg. Kalanchoe. Phytotaxa 2022, 550(2), 208-212. https://doi.org/10.11646/phytotaxa.550.2.10 |
[41] | Smith, G. F. Kalanchoe sect. Streptanthae (Crassulaceae subfam. Kalanchooideae), a new two-species section in K. subg. Bryophyllum. Phytotaxa 2023, 609(4), 273-281. https://doi.org/10.11646/phytotaxa.609.4.3 |
[42] | Smith, G. F. Publication of Kalanchoe sect. Trichanthae (Crassulaceae subfam. Kalanchooideae), a new status for K. [infragen. Unranked] Trichanthae of Alwin Berger (1930), in K. subg. Calophygia (2006 as amended in 2023). Phytotaxa 2023, 616(2), 176-182. https://doi.org/10.11646/phytotaxa.616.2.7 |
[43] | Smith, G. F. The unranked infrageneric names validly published in Kalanchoe (Crassulaceae subfam. Kalanchooideae) by Alwin Berger in 1930 revisited, with publication of five names at the rank of section and one at the rank of subgenus. Phytotaxa 2024, 635(4), 276-286. https://doi.org/10.11646/phytotaxa.635.4.2 |
[44] | Smith, G. F. Expanding and subdividing the southern and south-tropical African Kalanchoe [subg. Kalanchoe] sect. Raveta (Crassulaceae subfam. Kalanchooideae): Description of K. [sect. Raveta] ser. Raveta, K. [sect. Raveta] ser. Longiflorae, and K. [sect. Raveta] ser. Rotundifoliae. Phytotaxa 2024, 655(1), 1-20. https://doi.org/10.11646/phytotaxa.655.1.1 |
[45] | Smith, G. F. An alternative subgeneric classification of Kalanchoe (Crassulaceae subfam. Cotyledonoideae) globally: A nomenclatural and taxonomic review, with a new status for K. subg. Pubescentes. Phytotaxa 2024, 665(3), 201-220. https://doi.org/10.11646/phytotaxa.665.3.3 |
[46] | Smith, G. F. Refinements to the infra-sectional classification of Kalanchoe [subg. Bryophyllum] sect. Invasores (Crassulaceae subfam. Cotyledonoideae) at the rank of series: Description of K. [subg. Bryophyllum sect. Invasores] ser. Invasores, K. [subg. Bryophyllum sect. Invasores] ser. Fruticosa, and K. [subg. Bryophyllum sect. Invasores] ser. Herbacea. Phytotaxa 2025, 698(4), 196-208. https://doi.org/10.11646/phytotaxa. 698.4.1 |
[47] | Smith, G. F. Kalanchoe sect. Rhombopilosae (Crassulaceae subfam. Kalanchooideae), a new section for K. rhombopilosa in the woody clade of K. subg. Kalanchoe. Haseltonia 2023, 30(1), 22-26. https://doi.org/10.2985/026.030.0104 |
[2, 38-47]
Kalanchoe nothoser. Frutisores Z. H. Feng, nothoser. nov. = Kalanchoe ser. Fruticosa Gideon F. Sm. × Kalanchoe ser. Invasores Gideon F. Sm.
Kalanchoe nothoser. Herilana Z. H. Feng, nothoser. nov. = Kalanchoe ser. Herbacea Gideon F. Sm. × Kalanchoe ser. Vilana Gideon F. Sm.
Kalanchoe nothoser. Hersores Z. H. Feng, nothoser. nov. = Kalanchoe ser. Herbacea Gideon F. Sm. × Kalanchoe ser. Invasores Gideon F. Sm.
Kalanchoe nothoser. Invalana Z. H. Feng, nothoser. nov. = Kalanchoe ser. Invasores Gideon F. Sm. × Kalanchoe ser. Vilana Gideon F. Sm.
Petrosedum Grulich
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[48] | Afferni, M. Classification of the species of the genus Petrosedum Grulich in series. Avonia 2022, 40(2), 140-149. |
[49] | Afferni, M. Classification of the species of the genus Petrosedum Grulich in series: Further observations and corrections. Avonia 2022, 40(4), 349. |
[2, 48, 49]
Remark: Afferni assigned the genotype
Petrosedum rupestre to
Petrosedum ser.
Reflexa, according to Art. 22.1 of
ICN [4] | Turland, N., Wiersema, J., Barrie, F., Greuter, W., Hawksworth, D., Herendeen, P., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T., McNeill, J., Monro, A., Prado, J., Price, M., and Smith, G. eds. (2018) International Code of Nomenclature for algae, fungi, and plants. Koeltz Botanical Books. |
[4]
, the type serie must be named as
Petrosedum ser.
Petrosedum.
Petrosedum nothoser. Petrorecta Z. H. Feng, nothoser. nov. = Petrosedum ser. Petrosedum × Petrosedum ser. Recta Afferni.
Petrosedum nothoser. Petudorecta Z. H. Feng, nothoser. nov. = Petrosedum ser. Petrosedum × Petrosedum ser. Pseudorecta Afferni.
Petrosedum nothoser. Reseudorecta Z. H. Feng, nothoser. nov. = Petrosedum ser. Pseudorecta Afferni × Petrosedum ser. Recta Afferni.
Dryopteridaceae
Dryopteris Adans.
[50] | Fraser-Jenkins, C. R. A classification of the genus Dryopteris (Pteridophyta: Dryopteridaceae). Bull. Brit. Mus. (Nat. Hist.), Bot. 1986, 14(3), 183-218. |
[51] | Tzvelev, N. N. De genere Dryopteris Adans. (Dryopteridaceae) in Europa orientali. Novosti Sist. Vyssh. Rast. 2003, 35, 7-20. |
[52] | Kuo, L.-Y., Chang, Y.-H., Glowienka, J. M. O., Amoroso, V. B., Dong, S.-Y., Kao, T.-T., Wang, C.-N., Chiou, W.-L. A revised framework of Dryopteris subg. Nothoperanema (Dryopteridaceae) inferred from phylogenetic evidence, with descriptions of two new sections. Syst. Bot. 2016, 41(3), 596-605. https://doi.org/10.1600/036364416X692334 |
[53] | Zhang, L.-B., Zhang, L. The inclusion of Acrophorus, Diacalpe, Nothoperanema, and Peranema in Dryopteris: The molecular phylogeny, systematics, and nomenclature of Dryopteris subg. Nothoperanema (Dryopteridaceae). Taxon 2012, 61(6), 1199-1216. https://doi.org/10.1002/tax.616003 |
[54] | Zhang, L.-B., Zhang, L., Dong, S.-Y., Sessa, E. B., Gao, X.-F., Ebihara, A. Molecular circumscription and major evolutionary lineages of the fern genus Dryopteris (Dryopteridaceae). BMC Ecol. Evol. 2012, 12(1), 180. https://doi.org/10.1186/1471-2148-12-180 |
[50-54]
Dryopteris nothosect. Aedium Z. H. Feng, nothosect. nov. = Dryopteris sect. Aemulae Fraser-Jenk. × Dryopteris subgen. Lophodium (Newman) Tzvelev.
Dryopteris nothosect. Aemotae Z. H. Feng, nothosect. nov. = Dryopteris sect. Aemulae Fraser-Jenk. × Dryopteris sect. Remotae Fraser-Jenk.
Dryopteris nothosect. Aepteris Z. H. Feng, nothosect. nov. = Dryopteris sect. Aemulae Fraser-Jenk. × Dryopteris sect. Dryopteris.
Dryopteris nothosect. Aevariae Z. H. Feng, nothosect. nov. = Dryopteris sect. Aemulae Fraser-Jenk. × Dryopteris sect. Erythrovariae H. Ito.
Dryopteris nothosect. Drillodium Z. H. Feng, nothosect. nov. = Dryopteris sect. Dryopteris × Dryopteris sect. Fibrillosae Ching × Dryopteris subgen. Lophodium (Newman) Tzvelev.
Dryopteris nothosect. Dryllidae Z. H. Feng, nothosect. nov. = Dryopteris sect. Dryopteris × Dryopteris sect. Pallidae Fraser-Jenk.
Dryopteris nothosect. Dryollosae Z. H. Feng, nothosect. nov. = Dryopteris sect. Dryopteris × Dryopteris sect. Fibrillosae Ching.
Dryopteris nothosect. Dryphodium (Z. H. Feng) Z. H. Feng, stat. nov. = Dryopteris sect. Dryopteris × Dryopteris subgen. Lophodium (Newman) Tzvelev. — Basionym: Dryopteris nothosubgen. Dryphodium Z. H. Feng, this publication.
Dryopteris nothosect. Erypallidae Z. H. Feng, nothosect. nov. = Dryopteris sect. Erythrovariae H. Ito × Dryopteris sect. Pallidae Fraser-Jenk.
Dryopteris nothosect. Erypandae Z. H. Feng, nothosect. nov. = Dryopteris sect. Erythrovariae H. Ito × Dryopteris sect. Pandae Fraser-Jenk.
Dryopteris nothosect. Fidae Z. H. Feng, nothosect. nov. = Dryopteris sect. Fibrillosae Ching × Dryopteris sect. Pallidae Fraser-Jenk.
Dryopteris nothosect. Fiphodium Z. H. Feng, nothosect. nov. = Dryopteris sect. Fibrillosae Ching × Dryopteris subgen. Lophodium (Newman) Tzvelev.
Dryopteris nothosect. Hiropteris Z. H. Feng, nothosect. nov. = Dryopteris sect. Hirtipedes Fraser-Jenk. × Dryopteris subgen. Pycnopteris (Moore) Ching.
Dryopteris nothosect. Hirtidae Z. H. Feng, nothosect. nov. = Dryopteris sect. Hirtipedes Fraser-Jenk. × Dryopteris sect. Pallidae Fraser-Jenk.
Dryopteris nothosect. Hirtodon Z. H. Feng, nothosect. nov. = Dryopteris sect. Diclisodon (T. Moore) C. Chr. × Dryopteris sect. Hirtipedes Fraser-Jenk.
Dryopteris nothosect. Hirtosae Z. H. Feng, nothosect. nov. = Dryopteris sect. Fibrillosae Ching × Dryopteris sect. Hirtipedes Fraser-Jenk.
Dryopteris nothosect. Lolidae Z. H. Feng, nothosect. nov. = Dryopteris subgen. Lophodium (Newman) Tzvelev × Dryopteris sect. Pallidae Fraser-Jenk.
Dryopteris nothosect. Lophatae Z. H. Feng, nothosect. nov. = Dryopteris subgen. Lophodium (Newman) Tzvelev × Dryopteris sect. Marginatae Fraser-Jenk.
Dryopteris nothosect. Margisodon Z. H. Feng, nothosect. nov. = Dryopteris sect. Diclisodon (T. Moore) C. Chr. × Dryopteris sect. Marginatae Fraser-Jenk.
Dryopteris nothosect. Pallindae Z. H. Feng, nothosect. nov. = Dryopteris sect. Pallidae Fraser-Jenk. × Dryopteris sect. Pandae Fraser-Jenk.
Dryopteris nothosect. Palopteris Z. H. Feng, nothosect. nov. = Dryopteris sect. Pallidae Fraser-Jenk. × Dryopteris subgen. Pycnopteris (Moore) Ching.
Dryopteris nothosect. Panphodium Z. H. Feng, nothosect. nov. = Dryopteris subgen. Lophodium (Newman) Tzvelev × Dryopteris sect. Pandae Fraser-Jenk.
Dryopteris nothosect. Panpteris Z. H. Feng, nothosect. nov. = Dryopteris sect. Dryopteris × Dryopteris sect. Pandae Fraser-Jenk.
Dryopteris nothosect. Remolosae Z. H. Feng, nothosect. nov. = Dryopteris sect. Fibrillosae Ching × Dryopteris sect. Remotae Fraser-Jenk.
Dryopteris nothosect. Remophodium Z. H. Feng, nothosect. nov. = Dryopteris subgen. Lophodium (Newman) Tzvelev × Dryopteris sect. Remotae Fraser-Jenk.
Dryopteris nothosect. Remopteris Z. H. Feng, nothosect. nov. = Dryopteris sect. Dryopteris × Dryopteris sect. Remotae Fraser-Jenk.
Dryopteris nothosect. Shiopsis Z. H. Feng, nothosect. nov. = Dryopteris sect. Dryopsis (Holttum & P. J. Edwards) Li Bing Zhang × Dryopteris sect. Shiehia L. Y. Kuo & Y. H. Chang.
Dryopteris nothosect. Vavariae Z. H. Feng, nothosect. nov. = Dryopteris sect. Erythrovariae H. Ito × Dryopteris sect. Variae Fraser-Jenk.
Dryopteris nothosubgen. Dryphodium Z. H. Feng, nothosubgen. nov. = Dryopteris subgen. Dryopteris × Dryopteris subgen. Lophodium (Newman) Tzvelev.
Dryopteris nothosubgen. Erythteris Z. H. Feng, nothosubgen. nov. = Dryopteris subgen. Dryopteris × Dryopteris subgen. Erythrovariae (H. Ito) Fraser-Jenk.
Dryopteris nothosubgen. Lophema Z. H. Feng, nothosubgen. nov. = Dryopteris subgen. Lophodium (Newman) Tzvelev × Dryopteris subgen. Nothoperanema Tagawa.
Dryopteris nothosubgen. Notheris Z. H. Feng, nothosubgen. nov. = Dryopteris subgen. Dryopteris × Dryopteris subgen. Nothoperanema Tagawa.
Dryopteris nothosubgen. Nothovariae Z. H. Feng, nothosubgen. nov. = Dryopteris subgen. Erythrovariae (H. Ito) Fraser-Jenk. × Dryopteris subgen. Nothoperanema Tagawa.
Dryopteris nothosubgen. Pycyopteris Z. H. Feng, nothosubgen. nov. = Dryopteris subgen. Dryopteris × Dryopteris subgen. Pycnopteris (Moore) Ching.
Heliconiaceae
Heliconia L.
[55] | Kress, W. J., Fér, T., Carlsen, M. M. Phylogenomics and a new classification of the tropical genus Heliconia L. (Monocots, Zingiberales, Heliconiaceae). PhytoKeys 2025, 251, 37-66. https://doi.org/10.3897/phytokeys.251.130409 |
[55]
Heliconia nothosect. Epiconia Z. H. Feng, nothosect. nov. = Heliconia sect. Episcopales L. Anderss. × Heliconia sect. Heliconia.
Heliconia nothosect. Epistrobus Z. H. Feng, nothosect. nov. = Heliconia sect. Episcopales L. Anderss. × Heliconia sect. Taeniostrobus Kuntze.
Heliconia nothosubgen. Helitrobus Z. H. Feng, nothosubgen. nov. = Heliconia subgen. Heliconia × Heliconia subgen. Taeniostrobus (Kuntze) Griggs.
Hydrangeaceae
Hydrangea Gronov.
[56] | De Smet, Y., Granados Mendoza, C., Wanke, S., Goetghebeur, P., Samain, M.-S. Molecular phylogenetics and new (infra) generic classification to alleviate polyphyly in tribe Hydrangeeae (Cornales: Hydrangeaceae). Taxon 2015, 64(4), 741-753. https://doi.org/10.12705/644.6 |
[57] | Yang, X.-D., Xue, T.-T., Gao, T.-G., Liang, Y.-F., Smets, E. F., Gadagkar, S. R., Yu, S.-X. An updated infrageneric classification based on phylogenomics and character evolution in Hydrangea (Hydrangeaceae). Taxon 2024, 73(2), 503-518. https://doi.org/10.1002/tax.13158 |
[56, 57]
Hydrangea nothosect. Dichophyllae Z. H. Feng, nothosect. nov. = Hydrangea sect. Dichroa (Lour.) Y. De Smet & Samain × Hydrangea sect. Macrophyllae (E. M. McClint.) Y. De Smet & Samain.
Hydrangea nothosect. Hirtophyllae Z. H. Feng, nothosect. nov. = Hydrangea sect. Hirtae Y. De Smet & Samain × Hydrangea sect. Macrophyllae (E. M. McClint.) Y. De Smet & Samain.
Hydrangea nothosect. Kawirtae Z. H. Feng, nothosect. nov. = Hydrangea sect. Hirtae Y. De Smet & Samain × Hydrangea sect. Kawagoeanae X. D. Yang & S. X. Yu.
Hypericaceae
Hypericum nothosect. Drosoricum Z. H. Feng, nothosect. nov. = Hypericum sect. Drosocarpium Spach × Hypericum sect. Hypericum.
Hypericum nothoser. Drosoricum (Z. H. Feng) Z. H. Feng, stat. nov. = Hypericum sect. Drosocarpium Spach × Hypericum ser. Hypericum. — Basionym: Hypericum nothosect. Drosoricum Z. H. Feng, this publication.
Hypericum nothosubsect. Aethiosepalum Z. H. Feng, nothosubsect. nov. = Hypericum subsect. Adenosepalum N. Robson × Hypericum subsect. Aethiopica N. Robson.
Hypericum nothosubsect. Drosoricum (Z. H. Feng) Z. H. Feng, stat. nov. = Hypericum sect. Drosocarpium Spach × Hypericum subsect. Hypericum. — Basionym: Hypericum nothosect. Drosoricum Z. H. Feng, this publication.
Iridaceae
Gladiolus L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[59] | Goldblatt, P., Manning, J. C. Iridaceae of southern Africa. Strelitzia 2020, 42, 1-1159. |
[2, 59]
Gladiolus nothosect. Homifolii Z. H. Feng, nothosect. nov. = Gladiolus sect. Homoglossum (Salisb.) Goldblatt & J. C. Manning × Gladiolus sect. Linearifolii (M. P. de Vos) Goldblatt & J. C. Manning.
Gladiolus nothoser. Trearifolii Z. H. Feng, nothoser. nov. = Gladiolus ser. Linearifolii Goldblatt & J. C. Manning × Gladiolus ser. Tristes Goldblatt & J. C. Manning.
Lamiaceae
Lavandula L.
[60] | Passalacqua, N. G., Tundis, R., Upson, T. M. A new species of Lavandula sect. Lavandula (Lamiaceae) and review of species boundaries in Lavandula angustifolia. Phytotaxa 2017, 292(2), 161-170. https://doi.org/10.11646/phytotaxa.292.2.3 |
[61] | Patzelt, A., Al Hinai, A. Studies in the Flora of Arabia: XXXIII. A new species of Lavandula (Lamiaceae) from Oman. Edinburgh J. Bot. 2019, 76(3), 345-357. https://doi.org/10.1017/S0960428619000118 |
[62] | Suárez-Cervera, M. A., Seoane-Camba, J. A. Taxonomía numérica de algunas especies de Lavandula L., basada en caracteres morfológicos, cariológicos y palinológicos. Anales Jard. Bot. Madrid 1986, 42(2), 395-409. |
[63] | Upson, T., Andrews, S. The genus Lavandula. Timber Press; 2004, pp. 442. |
[60-63]
Lavandula nothosect. Laventatae Z. H. Feng, nothosect. nov. = Lavandula sect. Dentatae Suárez-Cerv. & Seoane-Camba × Lavandula sect. Lavandula.
Origanum L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[64] | Ietswaart, J. H. A Taxonomic Revision of the Genus Origanum (Labiatae). Leiden Botanical Series. No. 4. Leiden, the Netherlands: Leiden University Press; 1980, pp. 161. |
[2, 64]
Origanum nothosect. Amilamentum Z. H. Feng, nothosect. nov. = Origanum sect. Amaracus (Gled.) Vogel × Origanum sect. Brevifilamentum Ietsw.
Origanum nothosect. Amolicon Z. H. Feng, nothosect. nov. = Origanum sect. Amaracus (Gled.) Vogel × Origanum sect. Anatolicon Ietsw.
Origanum nothosect. Anatorana Z. H. Feng, nothosect. nov. = Origanum sect. Anatolicon Ietsw. × Origanum sect. Majorana (Mill.) Benth.
Origanum nothosect. Anatoriganum Z. H. Feng, nothosect. nov. = Origanum sect. Anatolicon Ietsw. × Origanum sect. Origanum.
Origanum nothosect. Brevicorolla Z. H. Feng, nothosect. nov. = Origanum sect. Brevifilamentum Ietsw. × Origanum sect. Prolaticorolla Ietsw.
Origanum nothosect. Brevifilicon Z. H. Feng, nothosect. nov. = Origanum sect. Anatolicon Ietsw. × Origanum sect. Brevifilamentum Ietsw.
Origanum nothosect. Eloticorolla Z. H. Feng, nothosect. nov. = Origanum sect. Elongatispica Ietsw. × Origanum sect. Prolaticorolla Ietsw.
Origanum nothosect. Longicorolla Z. H. Feng, nothosect. nov. = Origanum sect. Longitubus Ietsw. × Origanum sect. Prolaticorolla Ietsw.
Origanum nothosect. Origalyx Z. H. Feng, nothosect. nov. = Origanum sect. Chilocalyx (Briq.) Ietsw. × Origanum sect. Origanum.
Origanum nothosect. Origanomajorana (Domin) Z. H. Feng, comb. & stat. nov. = Origanum sect. Majorana (Mill.) Benth. × Origanum sect. Origanum. — Basionym: × Origanomajorana Domin, Preslia 13-15: 197. 1935.
Origanum nothosect. Orimentum Z. H. Feng, nothosect. nov. = Origanum sect. Brevifilamentum Ietsw. × Origanum sect. Origanum.
Origanum nothosect. Oriracus Z. H. Feng, nothosect. nov. = Origanum sect. Amaracus (Gled.) Vogel × Origanum sect. Origanum.
Origanum nothosect. Protolicon Z. H. Feng, nothosect. nov. = Origanum sect. Anatolicon Ietsw. × Origanum sect. Prolaticorolla Ietsw.
Teucrium nothosect. Chamaepolium Z. H. Feng, nothosect. nov. = Teucrium sect. Chamaedrys (Mill.) Schreb. × Teucrium sect. Polium (Mill.) Schreb.
Teucrium nothosect. Monpolium Z. H. Feng, nothosect. nov. = Teucrium sect. Montana Lázaro Ibiza × Teucrium sect. Polium (Mill.) Schreb.
Teucrium nothosect. Stachyodonia Z. H. Feng, nothosect. nov. = Teucrium sect. Scorodonia (Hill) Schreb. × Teucrium sect. Stachyobotrys Benth.
Teucrium nothosubsect. Canchamaedrys Z. H. Feng, nothosubsect. nov. = Teucrium subsect. Cana Melnikov × Teucrium subsect. Chamaedrys (Mill.) Kästner.
Teucrium nothosubsect. Chamaepolium (Z. H. Feng) Z. H. Feng, stat. nov. = Teucrium subsect. Chamaedrys (Mill.) Kästner × Teucrium subsect. Polium (Mill.) Kästner. — Basionym: Teucrium nothosect. Chamaepolium Z. H. Feng, this publication.
Teucrium nothosubsect. Monfolia Z. H. Feng, nothosubsect. nov. = Teucrium sect. Montana Lázaro Ibiza × Teucrium subsect. Rotundifolia M. E. Cohen.
Teucrium nothosubsect. Montolium Z. H. Feng, nothosubsect. nov. = Teucrium sect. Montana Lázaro Ibiza × Teucrium subsect. Polium (Mill.) Kästner.
Teucrium nothosubsect. Polifolia Z. H. Feng, nothosubsect. nov. = Teucrium subsect. Polium (Mill.) Kästner × Teucrium subsect. Rotundifolia M. E. Cohen.
Teucrium nothosubsect. Pomilum Z. H. Feng, nothosubsect. nov. = Teucrium subsect. Polium (Mill.) Kästner × Teucrium subsect. Pumilum Rivas Mart.
Teucrium nothosubsect. Popilosa Z. H. Feng, nothosubsect. nov. = Teucrium subsect. Polium (Mill.) Kästner × Teucrium subsect. Simplicipilosa S. Puech.
Teucrium nothosubsect. Pupilosa Z. H. Feng, nothosubsect. nov. = Teucrium subsect. Pumilum Rivas Mart. × Teucrium subsect. Simplicipilosa S. Puech.
Teucrium nothosubsect. Rotundipilosa Z. H. Feng, nothosubsect. nov. = Teucrium subsect. Rotundifolia M. E. Cohen × Teucrium subsect. Simplicipilosa S. Puech.
Lentibulariaceae
Pinguicula L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[66] | Fleischmann, A. On the infrageneric classification of Pinguicula. Carniv. Pl. Newslett. 2021, 50(4), 174-188. https://doi.org/10.55360/cpn504.af487 |
[67] | Fleischmann, A., Roccia, Systematics and evolution of Lentibulariaceae: I. Pinguicula. In Carnivorous Plants: Physiology, Ecology, and Evolution, Oxford University Press; 2018, pp. 70-80. https://doi.org/10.1093/oso/9780198779841.003.0006 |
[2, 66, 67]
Pinguicula nothosect. Micula Z. H. Feng, nothosect. nov. = Pinguicula sect. Micranthus Casper × Pinguicula sect. Pinguicula.
Pinguicula nothosubgen. Pinoceras Z. H. Feng, nothosubgen. nov. = Pinguicula subgen. Pinguicula × Pinguicula subgen. Temnoceras Barnhart.
Marattiaceae
Danaea Sm.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[68] | Presl, K. B. Supplementum tentaminis pteridographiae, continens genera et species ordinum dictorum marattiaceae, ophioglossaceae, osmundaceae, schizaeaceae et lygodiaceae. E typographia Cass. reg. Aulica filiorum Amadei Haase, Pragae; 1845, pp. 570. https://doi.org/10.5962/bhl.title.82188 |
[69] | Keskiniva, V., Tuomisto, H., Lehtonen, S. Danaea (Marattiaceae) keeps diversifying, part 2: Phylogeny and identification key for 81 taxa. Willdenowia 2024, 53(3), 229-255. https://doi.org/10.3372/wi.53.53304 |
[2, 68, 69]
Danaea nothosubgen. Dalodanaea Z. H. Feng, nothosubgen. nov. = Danaea subgen. Danaea × Danaea subgen. Holodanaea C. Presl.
Danaea nothosubgen. Darodanaea Z. H. Feng, nothosubgen. nov. = Danaea subgen. Arthrodanaea C. Presl × Danaea subgen. Danaea.
Musaceae
Musa L.
[70] | Li, L.-F., Häkkinen, M., Yuan, Y.-M., Hao, G., Ge, X.-J. Molecular phylogeny and systematics of the banana family (Musaceae) inferred from multiple nuclear and chloroplast DNA fragments, with a special reference to the genus Musa. Molec. Phylogen. Evol. 2010, 57, 1-10. https://doi.org/10.1016/j.ympev.2010.06.021 |
[70]
Musa nothosect. Mulimusa Z. H. Feng, nothosect. nov. = Musa sect. Australimusa Cheesman × Musa sect. Musa.
Nymphaeaceae
Nymphaea L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[17] | Conard, H. S. The waterlilies: A monograph of the genus Nymphaea. Carnegie Institution of Washington; 1905, pp. 279. https://doi.org/10.5962/bhl.title.23557 |
[18] | Löhne, C., Yoo, M.-J., Borsch, T., Wiersema, J., Wilde, V., Bell, C. D., Barthlott, W., Soltis, D. E., Soltis, P. S. Biogeography of Nymphaeales: Extant patterns and historical events. Taxon 2008, 57(4), 1123-1146. https://doi.org/10.1002/tax.574008 |
[2, 17, 18]
Nymphaea nothosect. Nymtha Zi Jun Li & Z. H. Feng, nothosect. nov. = Nymphaea sect. Nymphaea × Nymphaea sect. Xanthantha (Casp.) Wiersema.
Nymphaea nothosubgen.
Aneceras Zi Jun Li & Z. H. Feng, nothosubgen. nov. =
Nymphaea subgen.
Anecphya (Casp.) Conard
×
Nymphaea subgen.
Brachyceras (Casp.) Conard. (
Figure 1)
Nymphaea nothosubgen.
Anelotos Zi Jun Li & Z. H. Feng, nothosubgen. nov. =
Nymphaea subgen.
Anecphya (Casp.) Conard
×
Nymphaea subgen.
Lotos (DC.) Conard. (
Figure 1)
Nymphaea nothosubgen.
Brachynymphaea Zi Jun Li & Z. H. Feng, nothosubgen. nov. =
Nymphaea subgen.
Brachyceras (Casp.) Conard
×
Nymphaea subgen.
Nymphaea. (
Figure 1)
Orchidaceae
Anacamptis Rich.
[71] | Kretzschmar, H., Eccarius, W., Dietrich, H. The Orchid Genera Anacamptis, Orchis and Neotinea: Phylogeny, Taxonomy, Morphology, Biology, Distribution, Ecology and Hybridisation (2. ed). EchinoMedia; 2007, pp. 544. |
[72] | Kuropatkin, V. V., Efimov, P. G. A review of the genera Anacamptis, Neotinea and Orchis s. str. (Orchidaceae) in the flora of Russia and adjacent countries, with considerations of subdivision of Orchis s. l. Bot. Zhurn. (Moscow & Leningrad) 2014, 99(5), 555-593. |
[71, 72]
Anacamptis nothosect. Anacranthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Anacamptis × Anacamptis sect. Andranthus (Schltr.) F. M. Vázquez. — Synonym: × Anacampaludorchis P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Anacamptis nothosect. Andorianthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Andranthus (Schltr.) F. M. Vázquez × Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov.
Anacamptis nothosect. Andraenanthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Andranthus (Schltr.) F. M. Vázquez × Anacamptis sect. Phalaenanthus (Schltr.) Kuropatkin & Efimov.
Anacamptis nothosect. Andriopanthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Andranthus (Schltr.) F. M. Vázquez × Anacamptis sect. Coriophorae (Parl.) P. Quentin × Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov.
Anacamptis nothosect. Andriophorae Z. H. Feng, nothosect. nov. = Anacamptis sect. Andranthus (Schltr.) F. M. Vázquez × Anacamptis sect. Coriophorae (Parl.) P. Quentin. — Synonym: × Anteriopaludorchis P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Anacamptis nothosect. Andryae Z. H. Feng, nothosect. nov. = Anacamptis sect. Andranthus (Schltr.) F. M. Vázquez × Anacamptis sect. Boryae H. Kretzschmar, Eccarius & H. Dietr.
Anacamptis nothosect. Anlaenanthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Anacamptis × Anacamptis sect. Phalaenanthus (Schltr.) Kuropatkin & Efimov. — Synonym: × Anacampteulenia P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Anacamptis nothosect. Anorianthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Anacamptis × Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov. — Synonym: × Anacampterorchis P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Anacamptis nothosect. Anoriophorae Z. H. Feng, nothosect. nov. = Anacamptis sect. Anacamptis × Anacamptis sect. Coriophorae (Parl.) P. Quentin. — Synonym: × Anteriocamptis P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Figure 1. Representative cultivars from three Nymphaea nothosubgenera: A-B. Nymphaea ‘New Poem of the Nile’ (Nymphaea nothosubgen. Aneceras), C-D. Nymphaea ‘Tiwa Watree’ (Nymphaea nothosubgen. Anelotos), E-F. Nymphaea ‘Detective Erika’, G-H. Nymphaea ‘Siam Pink 1’ (Nymphaea nothosubgen. Brachynymphaea). All photos by Zi Jun Li.
Anacamptis nothosect. Boraenanthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Boryae H. Kretzschmar, Eccarius & H. Dietr. × Anacamptis sect. Phalaenanthus (Schltr.) Kuropatkin & Efimov.
Anacamptis nothosect. Borianthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Boryae H. Kretzschmar, Eccarius & H. Dietr. × Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov.
Anacamptis nothosect. Borophorae Z. H. Feng, nothosect. nov. = Anacamptis sect. Boryae H. Kretzschmar, Eccarius & H. Dietr. × Anacamptis sect. Coriophorae (Parl.) P. Quentin.
Anacamptis nothosect. Colaenanthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Coriophorae (Parl.) P. Quentin × Anacamptis sect. Phalaenanthus (Schltr.) Kuropatkin & Efimov. — Synonym: × Anteriomeulenia P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Anacamptis nothosect. Corianthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Coriophorae (Parl.) P. Quentin × Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov. — Synonym: × Anterioherorchis P. Delforge, Naturalistes Belges 90(Orchid. 22): 30. 2009.
Anacamptis nothosect. Moraccatae Z. H. Feng, nothosect. nov. = Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov × Anacamptis sect. Saccatae (Rchb. f.) P. Quentin.
Anacamptis nothosect. Moraenanthus Z. H. Feng, nothosect. nov. = Anacamptis sect. Morianthus (Schltr.) Kuropatkin & Efimov × Anacamptis sect. Phalaenanthus (Schltr.) Kuropatkin & Efimov.
Anacamptis nothosect. Phalaccatae Z. H. Feng, nothosect. nov. = Anacamptis sect. Phalaenanthus (Schltr.) Kuropatkin & Efimov × Anacamptis sect. Saccatae (Rchb. f.) P. Quentin.
Calanthe R. Br.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[6] | Clayton, D., Cribb, P. The genus Calanthe. Natural History Publications (Borneo) in association with Royal Botanic Gardens, Kew, Kota Kinabalu; 2013, pp. 413. |
[7] | Zhai, J.-W., Zhang, G.-Q., Li, L., Wang, M., Chen, L.-J., Chung, S.-W., Rodríguez, F. J., Francisco-Ortega, J., Lan, S.-R., Xing, F.-W., Liu, Z.-J. A new phylogenetic analysis sheds new light on the relationships in the Calanthe alliance (Orchidaceae) in China. Molec. Phylogen. Evol. 2014, 77, 216-222. https://doi.org/10.1016/j.ympev.2014.04.005 |
[8] | Chase, M. W., Christenhusz, M. J. M., Schuiteman, A. Expansion of Calanthe to include the species of Cephalantheropsis, Gastrorchis and Phaius (Collabieae; Orchidaceae). Phytotaxa 2020, 472(2), 159-168. https://doi.org/10.11646/phytotaxa.472.2.6 |
[9] | Chase, M. W., Christenhusz, M. J. M., Schuiteman, A. (2782) Proposal to conserve Calanthe, nom. cons., against the additional names Phaius, Cyanorkis, and Gastorkis (Orchidaceae, Collabieae). Taxon 2020, 69(6), 1364-1365. https://doi.org/10.1002/tax.12396 |
[10] | Wilson, K. L. Report of the General Committee: 26. Taxon 2023, 72(4), 905-907. https://doi.org/10.1002/tax.12994 |
[11] | Applequist, W. L. Report of the Nomenclature Committee for Vascular Plants: 73. Taxon 2023, 72(1), 179-204. https://doi.org/10.1002/tax.12871 |
[2, 6-11]
Calanthe nothosect. Triverna Z. H. Feng, nothosect. nov. = Calanthe sect. Tricarinata J. W. Zhai, Z. J. Liu & F. W. Xing × Calanthe sect. Verna Kakadzu.
Calanthe nothosubgen. Gastrocalanthe (A. D. Hawkes) Z. H. Feng, comb. & stat. nov. = Calanthe subgen. Calanthe × Calanthe subgen. Gastrorchis (Schltr.) Z. H. Feng. — Basionym: × Gastrocalanthe A. D. Hawkes, Orchid J. 1: 245. 1952.
Calanthe nothosubgen. Gastrophaius (A. D. Hawkes) Z. H. Feng, comb. & stat. nov. = Calanthe subgen. Gastrorchis (Schltr.) Z. H. Feng × Calanthe subgen. Phaius (Lour.) Z. H. Feng. — Basionym: × Gastrophaius A. D. Hawkes, Orchid J. 1: 245. 1952.
Calanthe nothosubgen. Phaiocalanthe (Rolfe) Z. H. Feng, comb. & stat. nov. = Calanthe subgen. Calanthe × Calanthe subgen. Phaius (Lour.) Z. H. Feng. — Basionym: × Phaiocalanthe Rolfe, J. Linn. Soc., Bot. 24: 168. 1887.
Calanthe nothosubgen. Phaiopreptanthe (Kerch.) Z. H. Feng, comb. & stat. nov. = Calanthe subgen. Phaius (Lour.) Z. H. Feng × Calanthe subgen. Preptanthe (Rchb. f.) Schltr. — Basionym: × Phaiopreptanthe Kerch., Le Liv. Orch.: 485. 1894.
Calanthe nothosubgen. Preptacalanthe (J. M. H. Shaw) Z. H. Feng, comb. & stat. nov. = Calanthe subgen. Calanthe × Calanthe subgen. Preptanthe (Rchb. f.) Schltr. — Basionym: × Preptacalanthe J. M. H. Shaw, Orchid Rev. Suppl., 121(1301): 14. 2013.
Cypripedium L.
[73] | Cribb, P. J. The genus Cypripedium. Timber Press; 1997, pp. 301. |
[74] | Eccarius, W. Die Orchideengattung Cypripedium: Phylogenie, Taxonomie, Morphologie, Biologie, Verbreitung, Ökologie und Hybridisation. EchinoMedia-Verl, Bürgel; 2009, pp. 384. |
[75] | Frosch, W., Cribb, P. J. Hardy Cypripedium: Species, hybrids and cultivation. Kew Publishing, Royal Botanic Gardens, Richmond, UK; 2012, pp. 156. |
[76] | Chen, S. C., Liu, Z.-J. Chen, L.-J. Li, L.-Q. The genus Cypripedium in China. Science Press, Beijing, China; 2013, pp. 317. |
[77] | Pérez-García, E. A., Mó, E. Die Cypripedioideae von Mesoamerika: Teil 1a Cypripedium. Orchidee (Hamburg) 2014, 65(6): 476-482. |
[78] | Szlachetko, D. L., Górniak, M., Kowalkowska, A. K., Kolanowska, M., Jurczak-Kurek, A., Archila Morales, F. The natural history of the genus Cypripedium (Orchidaceae). Pl. Biosystems 2021, 155(4), 772-796. https://doi.org/10.1080/11263504.2020.1785963 |
[73-78]
Cypripedium nothosect. Cypetala Z. H. Feng, nothosect. nov. = Cypripedium sect. Cypripedium × Cypripedium sect. Obtusipetala (Pfitzer) P. J. Cribb.
Cypripedium nothosubsect. Cypantha Z. H. Feng, nothosubsect. nov. = Cypripedium subsect. Cypripedium × Cypripedium subsect. Macrantha (Kraenzl.) P. J. Cribb.
Cypripedium nothosubsect. Obtupedium Z. H. Feng, nothosubsect. nov. = Cypripedium subsect. Cypripedium × Cypripedium sect. Obtusipetala (Pfitzer) P. J. Cribb.
Dactylorhiza Neck. ex Nevski
Dactylorhiza nothosect. Anculatae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Angorhiza Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Dactylorhiza.
Dactylorhiza nothosect. Aristoglossum Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Aristatae Aver. × Dactylorhiza sect. Coeloglossum (Hartm.) Efimov.
Dactylorhiza nothosect. Coelofoliae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Coeloglossum (Hartm.) Efimov.
Dactylorhiza nothosect. Coelolatae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Coeloglossum (Hartm.) Efimov × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Dactycinae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Dactylorhiza × Dactylorhiza sect. Sambucinae (Parl.) Smoljan.
Dactylorhiza nothosect. Dactyloglossum (P. F. Hunt & Summerh.) Z. H. Feng, comb. & stat. nov. = Dactylorhiza sect. Coeloglossum (Hartm.) Efimov × Dactylorhiza sect. Dactylorhiza. — Basionym: × Dactyloglossum P. F. Hunt & Summerh., Watsonia 6: 132. 1965.
Dactylorhiza nothosect. Daculacinae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Dactylorhiza × Dactylorhiza sect. Maculatae (Parl.) Smoljan. × Dactylorhiza sect. Sambucinae (Parl.) Smoljan.
Dactylorhiza nothosect. Ibefoliae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Iberanthus (Schltr.) Smoljan.
Dactylorhiza nothosect. Iberatae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Iberanthus (Schltr.) Smoljan. × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Iberorhiza Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Dactylorhiza × Dactylorhiza sect. Iberanthus (Schltr.) Smoljan.
Dactylorhiza nothosect. Macoglossum Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Coeloglossum (Hartm.) Efimov × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Macucinae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Maculatae (Parl.) Smoljan. × Dactylorhiza sect. Sambucinae (Parl.) Smoljan.
Dactylorhiza nothosect. Macufoliae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Macurhiza Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Dactylorhiza × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Macustirhiza Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Dactylorhiza × Dactylorhiza sect. Maculatae (Parl.) Smoljan.
Dactylorhiza nothosect. Samboglossum Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Coeloglossum (Hartm.) Efimov × Dactylorhiza sect. Sambucinae (Parl.) Smoljan.
Dactylorhiza nothosect. Samtifoliae Z. H. Feng, nothosect. nov. = Dactylorhiza sect. Angustifoliae (Verm.) Smoljan. × Dactylorhiza sect. Sambucinae (Parl.) Smoljan.
Gymnadenia R. Br.
[80] | Eccarius, W. Die Orchideengattung Gymnadenia: Mit einem Exkurs zur Gattung Pseudorchis: Systematik, Taxonomie, Morphologie, Biologie, Verbreitung, Ökologie und Hybridisation. Selbstverlag der Verfasser, Eisenach; 2022, pp. 336. |
[81] | Oddone, L., Andreoli, M. V., Casabianca, A. New genus subdivisions and taxonomic revision of intergeneric hybrids in genera Dactylorhiza, Gymnadenia, Pseudorchis and in nothogenera ×Pseudorhiza, ×Pseudadenia and ×Dactylodenia (Orchidaceae). GIROS Orch. Spont. Eur. 2016, 59(2): 347-377. |
[82] | Bateman, R. M. Phenotypic versus genotypic disparity in the Eurasian orchid genus Gymnadenia: Exploring the limits of phylogeny reconstruction. Syst. Biodivers. 2021, 19(4), 400-422. https://doi.org/10.1080/14772000.2021.1877845 |
[80-82]
Gymnadenia nothosect. Balcatiana Z. H. Feng, nothosect. nov. = Gymnadenia sect. Balcanica Eccarius × Gymnadenia sect. Praegnatiana Eccarius.
Gymnadenia nothosect. Botenia Z. H. Feng, nothosect. nov. = Gymnadenia sect. Botanicana Eccarius × Gymnadenia sect. Gymnadenia.
Gymnadenia nothosect. Gygnatiana Z. H. Feng, nothosect. nov. = Gymnadenia sect. Gymnadenia × Gymnadenia sect. Praegnatiana Eccarius.
Gymnadenia nothosect. Gymneiniana Z. H. Feng, nothosect. nov. = Gymnadenia sect. Gymnadenia × Gymnadenia sect. Wettsteiniana Eccarius.
Gymnadenia nothosect. Gymnigratiana Z. H. Feng, nothosect. nov. = Gymnadenia sect. Gymnadenia × Gymnadenia sect. Nigritella (Rich.) O. Gerbaud & P. Quentin × Gymnadenia sect. Praegnatiana Eccarius.
Gymnadenia nothosect. Gymnigritella (E. G. Camus) Z. H. Feng, comb. & stat. nov. = Gymnadenia sect. Gymnadenia × Gymnadenia sect. Nigritella (Rich.) O. Gerbaud & P. Quentin. — Basionym: × Gymnigritella E. G. Camus, J. Bot. (Morot) 6: 484. 1892.
Gymnadenia nothosect. Nigratiana Z. H. Feng, nothosect. nov. = Gymnadenia sect. Nigritella (Rich.) O. Gerbaud & P. Quentin × Gymnadenia sect. Praegnatiana Eccarius.
Gymnadenia nothosect. Praegeiniana Z. H. Feng, nothosect. nov. = Gymnadenia sect. Praegnatiana Eccarius × Gymnadenia sect. Wettsteiniana Eccarius.
Himantoglossum Spreng.
[83] | Bateman, R. M., Molnár V., A., & Sramkó, G. In situ morphometric survey elucidates the evolutionary systematics of the Eurasian Himantoglossum clade (Orchidaceae: Orchidinae). PeerJ 2017, 5, e2893. https://doi.org/10.7717/peerj.2893 |
[83]
Himantoglossum nothosubgen. Comptoglossum (Karatzas) Z. H. Feng, comb. & stat. nov. = Himantoglossum subgen. Comperia (K. Koch) R. M. Bateman, Molnar & Sramkó × Himantoglossum subgen. Himantoglossum. — Basionym: × Comptoglossum Karatzas, J. Eur. Orch. 36(4): 954. 2004.
Himantoglossum nothosect. Comprinum Z. H. Feng, nothosect. nov. = Himantoglossum subgen. Comperia (K. Koch) R. M. Bateman, Molnar & Sramkó × Himantoglossum sect. Caprinum R. M. Bateman, Molnar & Sramkó.
Ophrys L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[84] | Hennecke, M. Matching molecular genetics and morphology in the genus Ophrys. GIROS Orch. Spont. Eur. 2016, 59(1), 5-34. |
[85] | Hennecke, M. Beiträge zur Gattung Ophrys. Verlag Manfred Hennecke, Remshalden; 2021, pp. 656. |
[2, 84, 85]
Ophrys nothosect. Apidiniferae Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Apiflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Fuciflorae Rchb. f.
Ophrys nothosect. Apiliflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Bombyliflorae Rchb. f.
Ophrys nothosect. Apimuelleri Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Bornmuelleri Hennecke.
Ophrys nothosect. Apiniferae Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Araniferae Rchb. f.
Ophrys nothosect. Apiraniflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f.
Ophrys nothosect. Apiraniophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Ophrys.
Ophrys nothosect. Apispeculum Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Speculum Hennecke.
Ophrys nothosect. Apophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Apiferae P. Quentin × Ophrys sect. Ophrys.
Ophrys nothosect. Aracidophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Pseudophrys Godfery.
Ophrys nothosect. Araneudophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Pseudophrys Godfery.
Ophrys nothosect. Araniciflerae Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Aranispeculum Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Speculum Hennecke.
Ophrys nothosect. Aranomiflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f.
Ophrys nothosect. Aranophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Ophrys.
Ophrys nothosect. Aranyliflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Bombyliflorae Rchb. f.
Ophrys nothosect. Arelleri Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Bornmuelleri Hennecke.
Ophrys nothosect. Ariflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f.
Ophrys nothosect. Arumbilicatae Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Umbilicatae P. Quentin.
Ophrys nothosect. Bombeculum Z. H. Feng, nothosect. nov. = Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Speculum Hennecke.
Ophrys nothosect. Bomdophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Pseudophrys Godfery.
Ophrys nothosect. Bomelleri Z. H. Feng, nothosect. nov. = Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Bornmuelleri Hennecke.
Ophrys nothosect. Bominiferae Z. H. Feng, nothosect. nov. = Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Boriflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Bornmuelleri Hennecke × Ophrys sect. Fuciflorae Rchb. f.
Ophrys nothosect. Bornicatae Z. H. Feng, nothosect. nov. = Ophrys sect. Bornmuelleri Hennecke × Ophrys sect. Umbilicatae P. Quentin.
Ophrys nothosect. Faniophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Ophrys.
Ophrys nothosect. Fuceudophrys Z. H. Feng, nothosect. nov. = Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Pseudophrys Godfery.
Ophrys nothosect. Fuciculum Z. H. Feng, nothosect. nov. = Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Speculum Hennecke.
Ophrys nothosect. Fuciliflorae (Z. H. Feng) Z. H. Feng, stat. nov. = Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Fuciflorae Rchb. f. — Basionym: Ophrys nothosubgen. Fuciliflorae Z. H. Feng, this publication.
Ophrys nothosect. Fuciniferae Z. H. Feng, nothosect. nov. = Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Fucophrys (Z. H. Feng) Z. H. Feng, stat. nov. = Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Ophrys. — Basionym: Ophrys nothosubgen. Fucophrys Z. H. Feng, this publication.
Ophrys nothosect. Pseudiniferae Z. H. Feng, nothosect. nov. = Ophrys sect. Pseudophrys Godfery × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Pseudolleri Z. H. Feng, nothosect. nov. = Ophrys sect. Bornmuelleri Hennecke × Ophrys sect. Pseudophrys Godfery.
Ophrys nothosect. Pseudospeculum Z. H. Feng, nothosect. nov. = Ophrys sect. Pseudophrys Godfery × Ophrys sect. Speculum Hennecke.
Ophrys nothosect. Speniferae Z. H. Feng, nothosect. nov. = Ophrys sect. Speculum Hennecke × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Spicatae Z. H. Feng, nothosect. nov. = Ophrys sect. Speculum Hennecke × Ophrys sect. Umbilicatae P. Quentin.
Ophrys nothosect. Tencatae Z. H. Feng, nothosect. nov. = Ophrys sect. Tenthrediniferae P. Quentin × Ophrys sect. Umbilicatae P. Quentin.
Ophrys nothosect. Tenthraniferae Z. H. Feng, nothosect. nov. = Ophrys sect. Araniferae Rchb. f. × Ophrys sect. Tenthrediniferae P. Quentin.
Ophrys nothosect. Umbiferae Z. H. Feng, nothosect. nov. = Ophrys sect. Fuciflorae Rchb. f. × Ophrys sect. Umbilicatae P. Quentin.
Ophrys nothosect. Umbiliflorae Z. H. Feng, nothosect. nov. = Ophrys sect. Bombyliflorae Rchb. f. × Ophrys sect. Umbilicatae P. Quentin.
Ophrys nothosubgen. Fuciliflorae Z. H. Feng, nothosubgen. nov. = Ophrys subgen. Bombyliflorae Hennecke × Ophrys subgen. Fuciflorae Hennecke & S. Munzinger.
Ophrys nothosubgen. Fucophrys Z. H. Feng, nothosubgen. nov. = Ophrys subgen. Fuciflorae Hennecke & S. Munzinger × Ophrys subgen. Ophrys.
Orchis L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[71] | Kretzschmar, H., Eccarius, W., Dietrich, H. The Orchid Genera Anacamptis, Orchis and Neotinea: Phylogeny, Taxonomy, Morphology, Biology, Distribution, Ecology and Hybridisation (2. ed). EchinoMedia; 2007, pp. 544. |
[72] | Kuropatkin, V. V., Efimov, P. G. A review of the genera Anacamptis, Neotinea and Orchis s. str. (Orchidaceae) in the flora of Russia and adjacent countries, with considerations of subdivision of Orchis s. l. Bot. Zhurn. (Moscow & Leningrad) 2014, 99(5), 555-593. |
[86] | Jin, W.-T., Schuiteman, A., Chase, M. W., Li, J.-W., Chung, S.-W., Hsu, T.-C., Jin, X.-H. Phylogenetics of subtribe Orchidinae s. l. (Orchidaceae; Orchidoideae) based on seven markers (plastid matK, psaB, rbcL, trnL-F, trnH-psba, and nuclear nrITS, Xdh): Implications for generic delimitation. BMC Pl. Biol. 2017, 17(1), 222. https://doi.org/10.1186/s12870-017-1160-x |
[2, 71, 72, 86]
Orchis nothosect. Anthroculae Z. H. Feng, nothosect. nov. = Orchis sect. Anthropophorae H. Kretzschmar, Eccarius & H. Dietr. × Orchis sect. Masculae Lindl.
Orchis nothosect. Anthrolicae Z. H. Feng, nothosect. nov. = Orchis sect. Anthropophorae H. Kretzschmar, Eccarius & H. Dietr. × Orchis sect. Italicae H. Kretzschmar, Eccarius & H. Dietr.
Orchis nothosect. Anthrorchis Z. H. Feng, nothosect. nov. = Orchis sect. Anthropophorae H. Kretzschmar, Eccarius & H. Dietr. × Orchis sect. Orchis.
Orchis nothosect. Italorchis Z. H. Feng, nothosect. nov. = Orchis sect. Italicae H. Kretzschmar, Eccarius & H. Dietr. × Orchis sect. Orchis.
Orchis nothosect. Mascalcare Z. H. Feng, nothosect. nov. = Orchis sect. Masculae Lindl. × Orchis sect. Robustocalcare Hautz.
Orchis nothosect. Masciales Z. H. Feng, nothosect. nov. = Orchis sect. Masculae Lindl. × Orchis sect. Provinciales Parl.
Orchis nothosect. Mascillae Z. H. Feng, nothosect. nov. = Orchis sect. Masculae Lindl. × Orchis sect. Pusillae Parl.
Orchis nothosect. Mascorchis (Z. H. Feng) Z. H. Feng, stat. nov. = Orchis sect. Masculae Lindl. × Orchis sect. Orchis. — Basionym: Orchis nothosubgen. Mascorchis Z. H. Feng, this publication.
Orchis nothosect. Provalcare Z. H. Feng, nothosect. nov. = Orchis sect. Provinciales Parl. × Orchis sect. Robustocalcare Hautz.
Orchis nothosect. Provillae Z. H. Feng, nothosect. nov. = Orchis sect. Provinciales Parl. × Orchis sect. Pusillae Parl.
Orchis nothosect. Pusilcare Z. H. Feng, nothosect. nov. = Orchis sect. Pusillae Parl. × Orchis sect. Robustocalcare Hautz.
Orchis nothosubgen. Mascorchis Z. H. Feng, nothosubgen. nov. = Orchis subgen. Masculae H. Kretzschmar, Eccarius & H. Dietr. × Orchis subgen. Orchis.
Paphiopedilum Pfitzer
[87] | Cribb, P. J. Classification of Paphiopedilum. Renziana 2011, 1, 6-29. |
[88] | Karasawa, K., Saito, K. A revision of the genus Paphiopedilum (Orchidaceae). Bull. Hiroshima Bot. Gard. 1982, 5, 1-69. |
[89] | Lee, Y.-I., Chung, M.-C., Sydara, K., Souliya, O., Aphay, S. L. Taxonomic placement of Paphiopedilum rungsuriyanum (Cypripedioideae; Orchidaceae) based on morphological, cytological and molecular analyses. Bot. Stud. (Taipei) 2017, 58(1), 16. https://doi.org/10.1186/s40529-017-0170-1 |
[90] | Liu, Z.-J., Chen, S.-C., Chen, L.-J., Lei, S. P. The genus Paphiopedilum in China. Science Press, Beijing, China; 2009, pp. 371. |
[91] | Tsai, C.-C., Liao, P.-C., Ko, Y.-Z., Chen, C.-H., Chiang, Y.-C. Phylogeny and Historical Biogeography of Paphiopedilum Pfitzer (Orchidaceae) Based on Nuclear and Plastid DNA. Frontiers Pl. Sci. (Online journal) 2020, 11, 126. https://doi.org/10.3389/fpls.2020.00126 |
[87-91]
Paphiopedilum nothosect. Coryobata Z. H. Feng, nothosect. nov. = Paphiopedilum sect. Barbata Kraenzl. × Paphiopedilum sect. Coryopedilum Pfitzer.
Paphiopedilum nothosect. Paphiobata Z. H. Feng, nothosect. nov. = Paphiopedilum sect. Barbata Kraenzl. × Paphiopedilum sect. Paphiopedilum.
Paphiopedilum nothosect. Paphipetalum Z. H. Feng, nothosect. nov. = Paphiopedilum sect. Paphiopedilum × Paphiopedilum sect. Pardalopetalum Hallier f. & Pfitzer.
Paphiopedilum nothosect. Pardadilum Z. H. Feng, nothosect. nov. = Paphiopedilum sect. Coryopedilum Pfitzer × Paphiopedilum sect. Pardalopetalum Hallier f. & Pfitzer.
Phragmipedium Rolfe
[92] | Braem, G. J. A re-evaluation of the infrageneric taxonomy of the genus Phragmipedium. Richardiana 2011, 12(1), 16-24. |
[93] | Cervera, F. A checklist of Phragpedium species. Orchid Digest 2020, 84(4), 195-227. |
[94] | Cervera, F. Phragmipedium taxonomy and the species concept: Who is correct, mother nature or us?. Orchid Digest 2020, 84(4), 244-252. |
[95] | Rodríguez Salas, M., Benavides Rainilla, J., R. Espinoza, J. Genetic relationships of Phragmipedium species (Orchidaceae) using amplified fragment lenght polymorphism (AFLP) analysis. Lankesteriana 2007, 7(3), 493-496. https://doi.org/10.15517/lank.v0i0.7931 |
[92-95]
Phragmipedium nothosect. Himantofolia Z. H. Feng, nothosect. nov. = Phragmipedium sect. Himantopetalum (Hallier f.) Garay × Phragmipedium sect. Lorifolia (Kraenzl.) Garay.
Phragmipedium nothosect. Loripedium Z. H. Feng, nothosect. nov. = Phragmipedium sect. Lorifolia (Kraenzl.) Garay × Phragmipedium sect. Phragmipedium.
Phragmipedium nothosect. Loripetalum Z. H. Feng, nothosect. nov. = Phragmipedium sect. Lorifolia (Kraenzl.) Garay × Phragmipedium sect. Platypetalum Pfitzer.
Vanda R. Br.
[96] | Gardiner, L. M., Cribb, P. J. Vanda. Renziana 2013, 3, 1-94. |
[97] | Gardiner, L. M., Kocyan, A., Motes, M., Roberts, D. L., Emerson, B. C. (2013). Molecular phylogenetics of Vanda and related genera (Orchidaceae). Bot. J. Linn. Soc. 2013, 173(4), 549-572. https://doi.org/10.1111/boj.12102 |
[98] | Motes, M. R. The natural genus Vanda: A monograph. Redland Press, Redland, Florida, USA; 2021, pp. 359. |
[99] | Motes, M. R., Gardiner, L. M., Roberts, D. L. Vanda section Dactylolobatae: A summary, two new species, and a key to identification. Orchid Digest 2015, 79(2), 98-104. |
[96-99]
Vanda nothosect. Deltollaria Z. H. Feng, nothosect. nov. = Vanda sect. Deltoglossa Christenson × Vanda sect. Lamellaria Lindl.
Vanda nothosect. Longivanda Z. H. Feng, nothosect. nov. = Vanda sect. Longicalcarata Christenson × Vanda sect. Vanda.
Vanda nothosect. Testacarata Z. H. Feng, nothosect. nov. = Vanda sect. Longicalcarata Christenson × Vanda sect. Testaceae L. M. Gardiner.
Papaveraceae
Corydalis DC.
[100] | Chen, J.-T., Lidén, M., Huang, X.-H., Zhang, L., Zhang, X.-J., Kuang, T.-H., Landis, J. B., Wang, D., Deng, T., Sun, H. An updated classification for the hyper-diverse genus Corydalis (Papaveraceae: Fumarioideae) based on phylogenomic and morphological evidence. J. Integr. Pl. Biol. 2023, 65(9), 2138-2156. https://doi.org/10.1111/jipb.13499 |
[100]
Corydalis nothosect. Corydacava Z. H. Feng, nothosect. nov. = Corydalis sect. Corydalis × Corydalis sect. Radix-cava Irmisch.
Meconopsis Vig.
[101] | Grey-Wilson, C. The genus Meconopsis: Blue poppies and their relatives. Kew Publishing, Royal Botanic Gardens, Kew, UK; 2014, pp. 408. |
[102] | Xiao, W., Simpson, B. B. A new infrageneric classification of Meconopsis (Papaveraceae) based on a well-supported molecular phylogeny. Syst. Bot. 2017, 42(2), 226-233. https://doi.org/10.1600/036364417X695466 |
[101, 102]
Meconopsis nothosect. Granopsis Z. H. Feng, nothosect. nov. = Meconopsis sect. Grandes Fedde × Meconopsis sect. Meconopsis.
Papaver L.
[103] | Elvebakk, A., Bjerke, J. W. Papaver recircumscribed: A review of neighbouring Papaveraceae genera, including Afropapaver nom. et stat. nov. and Oreomecon, a large, Arctic-Alpine genus. PhytoKeys 2024, 248, 105-188. https://doi.org/10.3897/phytokeys.248.121011 |
[103]
Papaver nothosect. Meconantha Z. H. Feng, nothosect. nov. = Papaver sect. Macrantha Elkan × Papaver sect. Meconidium Spach.
Papaver nothosect. Papadium Z. H. Feng, nothosect. nov. = Papaver sect. Papaver × Papaver sect. Rhoeadium Spach.
Plantaginaceae
Antirrhinum L.
[104] | Otero, A., Fernández-Mazuecos, M., Vargas, P. Evolution in the Model Genus Antirrhinum Based on Phylogenomics of Topotypic Material. Frontiers Pl. Sci. (Online journal) 2021, 12, 631178. https://doi.org/10.389/fpls.2021.631178 |
[105] | Fernández Casas, F. J. De Antirrhinis notulae. Fontqueria 1998, 48, 195-202. |
[104, 105]
Antirrhinum nothosect. Streptorrhinum Z. H. Feng, nothosect. nov. = Antirrhinum sect. Antirrhinum × Antirrhinum sect. Streptosepalum (Rothm.) Fern. Casas.
Plantago L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[106] | Mower, J. P., Guo, W.-H., Partha, R., Fan, W.-S., Levsen, N., Wolff, K., Nugent, J. M., Pabón-Mora, N., González, F. Plastomes from tribe Plantagineae (Plantaginaceae) reveal infrageneric structural synapormorphies and localized hypermutation for Plantago and functional loss of ndh genes from Littorella. Molec. Phylogen. Evol. 2021, 162, 107217. https://doi.org/10.1016/j.ympev.2021.107217 |
[2, 106]
Plantago nothosect. Lamprotago Z. H. Feng, nothosect. nov. = Plantago sect. Lamprosantha Decne. × Plantago sect. Plantago.
Plantago nothosect. Lanceisantha Z. H. Feng, nothosect. nov. = Plantago sect. Lamprosantha Decne. × Plantago sect. Lanceifolia Barnéoud.
Plantago nothosubgen. Psyllago Z. H. Feng, nothosubgen. nov. = Plantago subgen. Plantago × Plantago subgen. Psyllium (Juss.) Harms & Reiche.
Plumbaginaceae
Limonium Mill.
[107] | Koutroumpa, K., Theodoridis, S., Warren, B. H., Jiménez, A., Celep, F., Doğan, M., Romeiras, M. M., Santos-Guerra, A., Fernández-Palacios, J. M., Caujapé-Castells, J., Moura, M., Menezes De Sequeira, M., Conti, E. An expanded molecular phylogeny of Plumbaginaceae, with emphasis on Limonium (sea lavenders): Taxonomic implications and biogeographic considerations. Ecol. Evol. 2018, 8(24), 12397-12424. https://doi.org/10.1002/ece3.4553 |
[108] | Malekmohammadi, M., Koutroumpa, K., Crespo, M. B., Domina, G., Korotkova, N., Akhani, H., Von Mering, S., Borsch, T., Berendsohn, W. G. A taxonomic backbone for the Plumbaginaceae (Caryophyllales). PhytoKeys 2024, 243, 67-103. https://doi.org/10.3897/phytokeys.243.122784 |
[107, 108]
Limonium nothosect. Ironium Z. H. Feng, nothosect. nov. = Limonium sect. Iranolimon M. Malekm., Akhani & Borsch × Limonium sect. Limonium.
Ranunculaceae
Helleborus L.
[109] | Long, L.-F., Yamada, K., Ochiai, M. A comprehensive review of the morphological and molecular taxonomy of the genus Helleborus (Ranunculaceae). Rev. Agric. Sci. 2023, 11, 106-120. https://doi.org/10.7831/ras.11.0_106 |
[110] | Mathew, B. Hellebores. Alpine Garden Society, London, UK; 1989, pp. 180. |
[111] | Meiners, J., Debener, T., Schweizer, G., Winkelmann, T. Analysis of the taxonomic subdivision within the genus Helleborus by nuclear DNA content and genome-wide DNA markers. Sci. Hort. (Amsterdam) 2011, 128(1), 38-47. https://doi.org/10.1016/j.scienta.2010.12.011 |
[112] | Werner, K., Ebel, F. Life history of the genus Hellehorus L. (Ranunculaceae). Flora, Morphol. Geobot. Ecophysiol. 1994, 189(2), 97-130. https://doi.org/10.1016/S0367-2530(17)30578-9 |
[109-112]
Helleborus nothosect. Chenoborus Z. H. Feng, nothosect. nov. = Helleborus sect. Chenopus Schiffn. × Helleborus sect. Helleborus.
Helleborus nothosect. Dicastrum Z. H. Feng, nothosect. nov. = Helleborus sect. Dicarpon Ulbr. × Helleborus sect. Helleborastrum Spach.
Helleborus nothosect. Grirus Z. H. Feng, nothosect. nov. = Helleborus sect. Griphopus Spach × Helleborus sect. Helleborus.
Helleborus nothosect. Hellarpon Z. H. Feng, nothosect. nov. = Helleborus sect. Dicarpon Ulbr. × Helleborus sect. Helleborus.
Helleborus nothosect. Hellastrum (Z. H. Feng) Z. H. Feng, stat. nov. = Helleborus sect. Helleborastrum Spach × Helleborus sect. Helleborus. — Basionym: Helleborus nothosubgen. Hellastrum Z. H. Feng, this publication.
Helleborus nothosect. Hellecarpus Z. H. Feng, nothosect. nov. = Helleborus sect. Helleborus × Helleborus sect. Syncarpus Schiffn.
Helleborus nothosubgen. Hellastrum Z. H. Feng, nothosubgen. nov. = Helleborus subgen. Helleborastrum (Spach) K. Werner & F. Ebel × Helleborus subgen. Helleborus.
Pulsatilla Mill.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[113] | Sramkó, G., Laczkó, L., Volkova, P. A., Bateman, R. M., Mlinarec, J. Evolutionary history of the Pasque-flowers (Pulsatilla, Ranunculaceae): Molecular phylogenetics, systematics and rDNA evolution. Molec. Phylogen. Evol. 2019, 135, 45-61. https://doi.org/10.1016/j.ympev.2019.02.015 |
[2, 113]
Pulsatilla nothosect. Preotilla (Z. H. Feng) Z. H. Feng, stat. nov. = Pulsatilla sect. Preonanthus (DC.) Spach. × Pulsatilla sect. Pulsatilla. — Basionym: Pulsatilla nothosubgen. Preotilla Z. H. Feng, this publication.
Pulsatilla nothosect. Semipulsatilla Z. H. Feng, nothosect. nov. = Pulsatilla sect. Pulsatilla × Pulsatilla sect. Semicampanaria Zämelis & Paegle.
Pulsatilla nothoser. Patanthus Z. H. Feng, nothoser. nov. = Pulsatilla ser. Patentes (Aichele & Schweg.) Juz. ex Tamura × Pulsatilla sect. Preonanthus (DC.) Spach.
Pulsatilla nothoser. Patensatilla Z. H. Feng, nothoser. nov. = Pulsatilla ser. Patentes (Aichele & Schweg.) Juz. ex Tamura × Pulsatilla ser. Pulsatilla.
Pulsatilla nothoser. Patenses Z. H. Feng, nothoser. nov. = Pulsatilla ser. Patentes (Aichele & Schweg.) Juz. ex Tamura × Pulsatilla ser. Pratenses (Aichele & Schweg.) Juz. ex Tamura.
Pulsatilla nothoser. Prananthus Z. H. Feng, nothoser. nov. = Pulsatilla ser. Pratenses (Aichele & Schweg.) Juz. ex Tamura × Pulsatilla sect. Preonanthus (DC.) Spach.
Pulsatilla nothoser. Pratensatilla Z. H. Feng, nothoser. nov. = Pulsatilla ser. Pratenses (Aichele & Schweg.) Juz. ex Tamura × Pulsatilla ser. Pulsatilla.
Pulsatilla nothoser. Preotilla (Z. H. Feng) Z. H. Feng, stat. nov. = Pulsatilla sect. Preonanthus (DC.) Spach. × Pulsatilla ser. Pulsatilla. — Basionym: Pulsatilla nothosubgen. Preotilla Z. H. Feng, this publication.
Pulsatilla nothosubgen. Preotilla Z. H. Feng, nothosubgen. nov. = Pulsatilla subgen. Preonanthus (DC.) Juz. × Pulsatilla subgen. Pulsatilla.
Rosaceae
Cliffortia L.
[114] | Whitehouse, C. M. A monograph on the genus Cliffortia. Strelitzia 2021, 43, 1-477. |
[114]
Cliffortia nothosect. Acetae Z. H. Feng, nothosect. nov. = Cliffortia sect. Acerosae C. Whitehouse × Cliffortia sect. Costatae Weim.
Cliffortia nothosect. Multifiviae Z. H. Feng, nothosect. nov. = Cliffortia sect. Multifidae C. Whitehouse × Cliffortia sect. Multinerviae DC.
Cliffortia nothosect. Multoides Z. H. Feng, nothosect. nov. = Cliffortia sect. Multinerviae DC. × Cliffortia sect. Ruboides C. Whitehouse.
Cliffortia nothosect. Rufidae Z. H. Feng, nothosect. nov. = Cliffortia sect. Multifidae C. Whitehouse × Cliffortia sect. Ruboides C. Whitehouse.
Cotoneaster Medik.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[115] | Fryer, J., Hylmö, B. Cotoneasters: A comprehensive guide to shrubs for flowers, fruit and foliage. Timber Press, Portland London, London, UK; 2009, pp. 344. |
[2, 115]
Cotoneaster nothosect. Chaenaster (Z. H. Feng) Z. H. Feng, stat. nov. = Cotoneaster sect. Chaenopetalum Koehne × Cotoneaster sect. Cotoneaster. — Basionym: Cotoneaster nothosubgen. Chaenaster Z. H. Feng, this publication.
Cotoneaster nothosect. Chaenoflori Z. H. Feng, nothosect. nov. = Cotoneaster sect. Chaenopetalum Koehne × Cotoneaster sect. Densiflori T. T. Yu.
Cotoneaster nothoser. Frinosi Z. H. Feng, nothoser. nov. = Cotoneaster ser. Frigidi T. T. Yu × Cotoneaster ser. Pannosi Flinck & B. Hylmö.
Cotoneaster nothoser. Concantes Z. H. Feng, nothoser. nov. = Cotoneaster ser. Conspicui G. Klotz × Cotoneaster ser. Radicantes G. Klotz.
Cotoneaster nothoser. Frifolii Z. H. Feng, nothoser. nov. = Cotoneaster ser. Frigidi T. T. Yu × Cotoneaster ser. Salicifolii T. T. Yu.
Cotoneaster nothoser. Racemaster Z. H. Feng, nothoser. nov. = Cotoneaster ser. Cotoneaster × Cotoneaster ser. Racemiflori (Pojark.) G. Klotz.
Cotoneaster nothoser. Tomentaster Z. H. Feng, nothoser. nov. = Cotoneaster ser. Cotoneaster × Cotoneaster ser. Tomentosi J. Fryer & B. Hylmö.
Cotoneaster nothosubgen. Chaenaster Z. H. Feng, nothosubgen. nov. = Cotoneaster subgen. Chaenopetalum (Koehne) G. Klotz × Cotoneaster subgen. Cotoneaster.
Crataegus L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[116] | Christensen, K. I. Revision of Crataegus sect. Crataegus and nothosect. Crataeguineae (Rosaceae-Maloideae) in the Old World. Syst. Bot. Monogr. 1992, 35, 1-199. https://doi.org/10.2307/25027810 |
[117] | Christensen, K. I., Zarrei, M., Kuzmina, M., Talent, N., Lin, C., Dickinson, T. A. Crataegus × ninae-celottiae and C. × cogswellii (Rosaceae, Maleae), two spontaneously formed intersectional nothospecies. PhytoKeys 2014, 36, 1-26. https://doi.org/10.3897/phytokeys.36.6784 |
[118] | Liston, A., Weitemier, K. A., Letelier, L., Podani, J., Zong, Y., Liu, L., Dickinson, T. A. Phylogeny of Crataegus (Rosaceae) based on 257 nuclear loci and chloroplast genomes: Evaluating the impact of hybridization. PeerJ 2021, 9, e12418. https://doi.org/10.7717/peerj.12418 |
[119] | Phipps, J. B. Crataegus: A nomenclator for sectional and serial names. Taxon 1983, 32(4), 598-604. https://doi.org/10.2307/1221729 |
[120] | Phipps, J. B. Hawthorns and medlars: Royal Horticultural Society Plant Collector Guide. Timber Press, Portland, USA; 2003, pp. 180. |
[121] | Phipps, J. B., Robertson, K. R., Smith, P. G., Rohrer, J. R. A checklist of the subfamily Maloideae (Rosaceae). Canad. J. Bot. 1990, 68(10), 2209-2269. https://doi.org/10.1139/b90-288 |
[122] | Ufimov, R. A. Notes on the genus Crataegus L. (Rosaceae). Novosti Sist. Vyssh. Rast. 2013, 44, 113-125. https://doi.org/10.31111/novitates/2013.44.113 |
[123] | Ufimov, R. A., Dickison, T. A. Infrageneric nomenclature adjustments in Crataegus L. (Maleae, Rosaceae). Phytologia 2020, 102(3), 177-199. |
[2, 116-123]
Crataegus nothosect. Brespilus (Z. H. Feng) Z. H. Feng, stat. nov. = Crataegus sect. Brevispinae (Beadle) C. K. Schneid. × Crataegus sect. Mespilus (L.) T. A. Dickinson & E. Y. Y. Lo. — Basionym: Crataegus nothosubgen. Brespilus Z. H. Feng, this publication.
Crataegus nothosect. Coccanthae Z. H. Feng, nothosect. nov. = Crataegus sect. Coccineae Loudon × Crataegus sect. Macracanthae Loudon.
Crataegus nothoser. Anofoliae Z. H. Feng, nothoser. nov. = Crataegus ser. Anomalae (Sarg. ex Eggl.) Rehder × Crataegus ser. Tenuifoliae (Sarg. ex Eggl.) Rehder.
Crataegus nothoser. Anosae Z. H. Feng, nothoser. nov. = Crataegus ser. Anomalae (Sarg. ex Eggl.) Rehder × Crataegus ser. Pruinosae Sarg. ex Rehder.
Crataegus nothoser. Anothae Z. H. Feng, nothoser. nov. = Crataegus ser. Anomalae (Sarg. ex Eggl.) Rehder × Crataegus ser. Macracanthae (Loudon) Rehder.
Crataegus nothoser. Apolles Z. H. Feng, nothoser. nov. = Crataegus ser. Apiifoliae (Loudon) Rehder × Crataegus ser. Molles (Sarg. ex C. K. Schneider) Rehder.
Crataegus nothoser. Coccifoliae Z. H. Feng, nothoser. nov. = Crataegus ser. Coccineae (Loudon) Rehder × Crataegus ser. Tenuifoliae (Sarg. ex Eggl.) Rehder.
Crataegus nothoser. Crataegrae Z. H. Feng, nothoser. nov. = Crataegus ser. Crataegus × Crataegus ser. Nigrae (Loudon) Russanov.
Crataegus nothoser. Crusanthae Z. H. Feng, nothoser. nov. = Crataegus ser. Crus-galli (Loudon) Rehder × Crataegus ser. Macracanthae (Loudon) Rehder.
Crataegus nothoser. Incanthae Z. H. Feng, nothoser. nov. = Crataegus ser. Intricatae Sarg. ex Rehder × Crataegus ser. Macracanthae (Loudon) Rehder.
Crataegus nothoser. Innosae Z. H. Feng, nothoser. nov. = Crataegus ser. Intricatae Sarg. ex Rehder × Crataegus ser. Pruinosae Sarg. ex Rehder.
Crataegus nothoser. Macoliae Z. H. Feng, nothoser. nov. = Crataegus ser. Macracanthae (Loudon) Rehder × Crataegus ser. Parvifoliae (Loudon) Rehder.
Crataegus nothoser. Manosae Z. H. Feng, nothoser. nov. = Crataegus ser. Macracanthae (Loudon) Rehder × Crataegus ser. Pruinosae Sarg. ex Rehder.
Crataegus nothoser. Mollides Z. H. Feng, nothoser. nov. = Crataegus ser. Molles (Sarg. ex C. K. Schneider) Rehder × Crataegus ser. Virides (Beadle ex C. K. Schneider) Rehder.
Crataegus nothoser. Potigynae Z. H. Feng, nothoser. nov. = Crataegus ser. Pentagynae (C. K. Schneid.) Russanov × Crataegus ser. Poticae Pojark.
Crataegus nothoser. Prudifoliae Z. H. Feng, nothoser. nov. = Crataegus ser. Pruinosae Sarg. ex Rehder × Crataegus ser. Rotundifoliae (Eggl. ex Eggl.) Rehder.
Crataegus nothoser. Pugalli Z. H. Feng, nothoser. nov. = Crataegus ser. Crus-galli (Loudon) Rehder × Crataegus ser. Punctatae (Loudon) Rehder.
Crataegus nothoser. Puncosae Z. H. Feng, nothoser. nov. = Crataegus ser. Pruinosae Sarg. ex Rehder × Crataegus ser. Punctatae (Loudon) Rehder.
Crataegus nothoser. Rocanthae Z. H. Feng, nothoser. nov. = Crataegus ser. Rotundifoliae (Eggl. ex Eggl.) Rehder × Crataegus ser. Macracanthae (Loudon) Rehder.
Crataegus nothoser. Ronuifoliae Z. H. Feng, nothoser. nov. = Crataegus ser. Rotundifoliae (Eggl. ex Eggl.) Rehder × Crataegus ser. Tenuifoliae (Sarg.) Rehder.
Crataegus nothoser. Tentae Z. H. Feng, nothoser. nov. = Crataegus ser. Punctatae (Loudon) Rehder × Crataegus ser. Tenuifoliae (Sarg. ex Eggl.) Rehder.
Crataegus nothosubgen. Ametaegus Z. H. Feng, nothosubgen. nov. = Crataegus subgen. Americanae El Gazzar × Crataegus subgen. Crataegus.
Crataegus nothosubgen. Brespilus Z. H. Feng, nothosubgen. nov. = Crataegus subgen. Brevispinae (Beadle) Ufimov & T. A. Dickinson × Crataegus subgen. Mespilus (L.) Ufimov & T. A. Dickinson.
Crataegus nothosubgen. Crataeguineae (K. I. Chr.) Z. H. Feng, stat. nov. = Crataegus subgen. Crataegus × Crataegus subgen. Sanguineae Ufimov. — Basionym: Crataegus nothosect. Crataeguineae K. I. Chr., Syst. Bot. Monogr. 35: 161. 1992.
Prunus L.
[2] | POWO. 2025. Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed 1 June 2025) |
[124] | Shi, S., Li, J., Sun, J., Yu, J., Zhou, S. Phylogeny and classification of Prunus sensu lato (Rosaceae). J. Integr. Pl. Biol. 2013, 55(11), 1069-1079. https://doi.org/10.1111/jipb.12095 |
[2, 124]
Prunus nothosect. Micropurus Z. H. Feng, nothosect. nov. = Prunus sect. Microcerasus (Spach) C. K. Schneid. × Prunus sect. Prunus.
Prunus nothosect. Prudalus Z. H. Feng, nothosect. nov. = Prunus sect. Amygdalus (L.) Benth. & Hook. f. × Prunus sect. Prunus.
Prunus nothosect. Pruniaca Z. H. Feng, nothosect. nov. = Prunus sect. Armeniaca (Scop.) Koch × Prunus sect. Prunus.
Pyrus L.
[121] | Phipps, J. B., Robertson, K. R., Smith, P. G., Rohrer, J. R. A checklist of the subfamily Maloideae (Rosaceae). Canad. J. Bot. 1990, 68(10), 2209-2269. https://doi.org/10.1139/b90-288 |
[125] | Jin, Z.-T., Ma, D.-K., Liu, G.-N., Hodel, R. G. J., Jiang, Y., Ge, B.-J., Liao, S., Duan, L., Ren, C., Xu, C., Wu, J., Liu, B.-B. Advancing Pyrus phylogeny: Deep genome skimming-based inference coupled with paralogy analysis yields a robust phylogenetic backbone and an updated infrageneric classification of the pear genus (Maleae, Rosaceae). Taxon 2024, 73(3), 784-799. https://doi.org/10.1002/tax.13163 |
[121, 125]
Pyrus nothosubgen. Pyshia Z. H. Feng, nothosubgen. nov. = Pyrus subgen. Pashia (Koehne) B. B. Liu × Pyrus subgen. Pyrus.
Rubus L.
[126] | Huang, T.-R., Chen, J.-H., Hummer, K. E., Alice, L. A., Wang, W.-H., He, Y., Yu, S.-X., Yang, M.-F., Chai, T.-Y., Zhu, X.-Y., Ma, L.-Q., Wang, H. Phylogeny of Rubus (Rosaceae): Integrating molecular and morphological evidence into an infrageneric revision. Taxon 2023, 72(2), 278-306. https://doi.org/10.1002/tax.12885 |
[127] | Reveal, J. L. Lexicon of infrageneric names in Rubus (Rosaceae: Rubeae). Kew Bull. 2014, 69(3)-9524, 1-18. https://doi.org/10.1007/s12225-014-9524-y |
[126, 127]
Rubus nothosubgen. Bachobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Batothamnus (Focke) E. H. L. Krause × Rubus subgen. Malachobatus (Focke) Fritsch.
Rubus nothosubgen. Bateobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Batothamnus (Focke) E. H. L. Krause × Rubus subgen. Idaeobatus Focke.
Rubus nothosubgen. Chalanobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Chamaerubus Kuntze × Rubus subgen. Melanobatus (Greene) House.
Rubus nothosubgen. Cynobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Cylactis (Raf.) Focke × Rubus subgen. Melanobatus (Greene) House.
Rubus nothosubgen. Idanobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Idaeobatus Focke × Rubus subgen. Melanobatus (Greene) House.
Rubus nothosubgen. Linchobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Lineati (Focke) T. R. Huang & X. Y. Zhu × Rubus subgen. Malachobatus (Focke) Fritsch.
Rubus nothosubgen. Malnobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Malachobatus (Focke) Fritsch × Rubus subgen. Melanobatus (Greene) House.
Rubus nothosubgen. Rubobatus Z. H. Feng, nothosubgen. nov. = Rubus subgen. Melanobatus (Greene) House × Rubus subgen. Rubus.
Saxifragaceae
Saxifraga L.
[128] | Tkach, N., Röser, M., Miehe, G., Muellner-Riehl, A. N., Ebersbach, J., Favre, A., Hoffmann, M. H. Molecular phylogenetics, morphology and a revised classification of the complex genus Saxifraga (Saxifragaceae). Taxon 2015, 64(6), 1159-1187. https://doi.org/10.12705/646.4 |
[128]
Saxifraga nothosect. Cotypera Z. H. Feng, nothosect. nov. = Saxifraga sect. Cotylea Tausch × Saxifraga sect. Gymnopera D. Don.
Saxifraga nothosect. Liguphyrion Z. H. Feng, nothosect. nov. = Saxifraga sect. Ligulatae Haw. × Saxifraga sect. Porphyrion Tausch.
Saxifraga nothosect. Ligusaxifraga Z. H. Feng, nothosect. nov. = Saxifraga sect. Ligulatae Haw. × Saxifraga sect. Saxifraga.
Saxifraga nothosect. Trachysaxifraga Z. H. Feng, nothosect. nov. = Saxifraga sect. Saxifraga × Saxifraga sect. Trachyphyllum (Gaudin) W. D. J. Koch.
Saxifraga nothosubsect. Androsaxifraga Z. H. Feng, nothosubsect. nov. = Saxifraga subsect. Androsaceae (Engl. & Irmsch.) Tkach, Röser & M. H. Hoffm. × Saxifraga subsect. Saxifraga.
Saxifraga nothosubsect. Androtylites Z. H. Feng, nothosubsect. nov. = Saxifraga subsect. Androsaceae (Engl. & Irmsch.) Tkach, Röser & M. H. Hoffm. × Saxifraga subsect. Tridactylites (Haw.) Gornall.
Saxifraga nothosubsect. Araxifraga Z. H. Feng, nothosubsect. nov. = Saxifraga subsect. Arachnoideae (Engl. & Irmsch.) Tkach, Röser & M. H. Hoffm. × Saxifraga subsect. Saxifraga.
Saxifraga nothosubsect. Kabrrosae Z. H. Feng, nothosubsect. nov. = Saxifraga subsect. Kabschia Rouy & E. G. Camus × Saxifraga subsect. Squarrosae (Engl. & Irmsch.) Tkach, Röser & M. H. Hoffm.
Saxifraga nothosubsect. Kabulatae Z. H. Feng, nothosubsect. nov. = Saxifraga sect. Ligulatae Haw. × Saxifraga subsect. Kabschia Rouy & E. G. Camus.
Saxifraga nothosubsect. Ligusaxifraga (Z. H. Feng) Z. H. Feng, stat. nov. = Saxifraga sect. Ligulatae Haw. × Saxifraga subsect. Saxifraga. — Basionym: Saxifraga nothosect. Ligusaxifraga Z. H. Feng, this publication.
Saxifraga nothosubsect. Murrosae Z. H. Feng, nothosubsect. nov. = Saxifraga subsect. Mutatae (Engl. & Irmsch.) Gornall × Saxifraga subsect. Squarrosae (Engl. & Irmsch.) Tkach, Röser & M. H. Hoffm.
Saxifraga nothosubsect. Mutalatae Z. H. Feng, nothosubsect. nov. = Saxifraga sect. Ligulatae Haw. × Saxifraga subsect. Mutatae (Engl. & Irmsch.) Gornall.
Saxifraga nothosubsect. Trachysaxifraga (Z. H. Feng) Z. H. Feng, stat. nov. = Saxifraga subsect. Saxifraga × Saxifraga sect. Trachyphyllum (Gaudin) W. D. J. Koch. — Basionym: Saxifraga nothosect. Trachysaxifraga Z. H. Feng, this publication.
Saxifraga nothosubsect. Trisaxifraga Z. H. Feng, nothosubsect. nov. = Saxifraga subsect. Saxifraga × Saxifraga subsect. Tridactylites (Haw.) Gornall.
Verbenaceae
Verbena L.
[129] | Nesom, G. L. Infrageneric classification of Verbena (Verbenaceae). Phytoneuron 2010, 2010-11, 1-15. |
[129]
Verbena nothoser. Bracelabrae Z. H. Feng, nothoser. nov. = Verbena ser. Bracteatae G. L. Nesom × Verbena ser. Candelabrae G. L. Nesom.
Verbena nothoser. Canbena Z. H. Feng, nothoser. nov. = Verbena ser. Candelabrae G. L. Nesom × Verbena ser. Verbena.
Verbena nothoser. Candachyae Z. H. Feng, nothoser. nov. = Verbena ser. Candelabrae G. L. Nesom × Verbena ser. Leptostachyae Schauer.
Verbena nothoser. Candelices Z. H. Feng, nothoser. nov. = Verbena ser. Candelabrae G. L. Nesom × Verbena ser. Simplices G. L. Nesom.
Verbena nothoser. Halabrae Z. H. Feng, nothoser. nov. = Verbena ser. Candelabrae G. L. Nesom × Verbena ser. Haleae G. L. Nesom.
Verbena nothoser. Leptacteatae Z. H. Feng, nothoser. nov. = Verbena ser. Bracteatae G. L. Nesom × Verbena ser. Leptostachyae Schauer.
Verbena nothoser. Leptices Z. H. Feng, nothoser. nov. = Verbena ser. Leptostachyae Schauer × Verbena ser. Simplices G. L. Nesom.
Verbena nothoser. Plibena Z. H. Feng, nothoser. nov. = Verbena ser. Plicatae G. L. Nesom × Verbena ser. Verbena.
Verbena nothoser. Plicteatae Z. H. Feng, nothoser. nov. = Verbena ser. Bracteatae G. L. Nesom × Verbena ser. Plicatae G. L. Nesom.
Violaceae
Viola L.
[130] | Marcussen, T., Ballard, H. E., Danihelka, J., Flores, A. R., Nicola, M. V., Watson, J. M. A revised phylogenetic classification for Viola (Violaceae). Plants (Basel) 2022, 11(17)-2224, 1-135. https://doi.org/10.3390/plants11172224 |
[130]
Viola nothosect. Plagioviola Z. H. Feng, nothosect. nov. = Viola sect. Plagiostigma Godr. × Viola sect. Viola.
Viola nothosubsect. Brebracteatae Z. H. Feng, nothosubsect. nov. = Viola subsect. Bracteolatae Kupffer × Viola subsect. Ebracteatae Kupffer.
Viola nothosubsect. Rostellares Z. H. Feng, nothosubsect. nov. = Viola subsect. Patellares (Boiss.) Rouy & Foucaud × Viola subsect. Rostratae (Kupffer) W. Becker.
Viola nothosubsect. Rostiola Z. H. Feng, nothosubsect. nov. = Viola subsect. Rostratae (Kupffer) W. Becker × Viola subsect. Viola.
3.3. New Nothogenera and Combinations of Pteridophytes and Spermatophytes
Apocynaceae
× Orbelia plowesii (P. V. Heath) Z. H. Feng, comb. nov. — Basionym: × Gonostapelia plowesii P. V. Heath, Calyx 1(1): 21. 1992.
Hybrid formula: Orbea maculata (N. E. Br.) L. C. Leach × Stapelia kwebensis N. E. Br.
Type: Zimbabwe. Mutema Pan, s. d., Plowes 3798 (holotype: PRE; possible isotype: SRGH).
× Tromostapelia bella (A. Berger) Z. H. Feng, comb. nov. — Basionym: Stapelia × bella A. Berger, Gard. Chron. ser. 3, 31: 137. 1902.
Hybrid formula: Stapelia deflexa Jacq. × Tromotriche revoluta (Masson) Haw.
Type: Provenance not given, 16 October 1901, A. Berger s. n. (holotype: K [barcode] K 000306220, digital image!).
Athyriaceae
Cornopteris bipinnata (K. Hori) Z. H. Feng, comb. nov. — Basionym: Athyrium bipinnatum K. Hori, PhytoKeys 148: 97. 2020.
Type: JAPAN. Shikoku: Kochi prefecture, Ochi town, Mt. Yokogura, 664 m, planted coniferous forest containing Cryptomeria japonica (Thunb. ex L. f.) D. Don, on soil, 29 June 2019, K. Hori 3277 (holotype: MAK [barcode] MAK 466762; isotype: MBK).
Cornopteris × decurrentiserrata (K. Hori) Z. H. Feng, comb. nov. — Basionym: Athyrium × decurrentiserratum K. Hori, Fern Gaz. 21(6): 262. 2021.
Hybrid formula: Cornopteris crenulatoserrulata (Makino) Nakai × Cornopteris decurrentialata (Hook.) Nakai.
Type: JAPAN. Honshu: Tokyo-to, Ome city, Koshizawa, 396 m, planted coniferous forest containing Cryptomeria japonica (Thunb. ex L. f.) D. Don, on soil, 4 June 2018, Hori 2981 (holotype: MAK [barcode] MAK 466750; isotype: MBK).
Cornopteris × pseudobipinnata (K. Hori) Z. H. Feng, comb. nov. — Basionym: Athyrium × pseudobipinnatum K. Hori, Fern Gaz. 21(6): 264. 2021.
Hybrid formula: Cornopteris bipinnata (K. Hori) Z. H. Feng × Cornopteris decurrentialata (Hook.) Nakai.
Type: JAPAN. Kyushu: Fukuoka pref., Itoshima city, Shiraito, 515 m, planted coniferous forest containing Cryptomeria japonica, on soil, 23 June 2019, Hori 3274 (holotype: MAK [barcode] MAK 466761; isotype: MBK).
Cornopteris × subchristenseniana (K. Hori) Z. H. Feng, comb. nov. — Basionym: Athyrium × subchristensenianum K. Hori, Fern Gaz. 21(6): 265. 2021.
Hybrid formula: Cornopteris crenulatoserrulata (Makino) Nakai × an unknown allotetraploid species.
Type: JAPAN. Honshu: Akita pref., Yurihonjo city, Hachinosawa, 180 m, planted coniferous forest containing Cryptomeria japonica (Thunb. ex L. f.) D. Don, on soil, 5 July 2019, Hori 3281 (holotype: MAK [barcode] MAK 466765; isotype: MBK).
Convolvulaceae
Convolvulus × turcicus nothosubsp. peshmenii (Aykurt & Sümbül) Z. H. Feng, comb. & stat. nov. — Basionym: Convolvulus × peshmenii Aykurt & Sümbül, Nordic J. Bot. 29(4): 409. 2011.
Hybrid formula: Convolvulus boissieri subsp. compactus (Boiss.) Stace × Convolvulus holosericeus subsp. macrocalycinus Hausskn. & Bornm.
Type: Turkey. southern Anatolia. Adana, Pozantı, 5-6 km from Pozantı to Adana, 945 m, 9 June 2007, C. Aykurt & N. Kemaloğlu 1495 (holotype: AKDU).
Lamiaceae
Salvia × hegelmaieri nothosubsp. accidentalis (Sánchez-Gómez & R. Morales) Z. H. Feng, comb. & stat. nov. — Basionym: Salvia × accidentalis Sánchez-Gómez & R. Morales, Anales Jard. Bot. Madrid 57(2): 421. 2000.
Hybrid formula: Salvia blancoana subsp. vellerea (Cuatrec.) W. Lippert × Salvia officinalis subsp. officinalis.
Type: SPAIN. Murcia. Moratalla, La Alberquilla, 10 May 1993, P. Sánchez Gómez s. n. (holotype: MA [barcode] MA 593325).
Orchidaceae
Calanthe × berryana (Sander) Z. H. Feng, comb. nov. — Basionym: × Phaiocalanthe berryana Sander, Gard. Chron. ser. 3, 18: 655. 1895.
Hybrid formula: Calanthe humblotii (Rchb. f.) M. W. Chase, Christenh. & Schuit. × Calanthe masuca (D. Don) Lindl.
Type: Not designated.
Calanthe × brandtiae (Sander) Z. H. Feng, comb. nov. — Basionym: × Phaiocalanthe brandtiae Sander, Gard. Chron. ser. 3, 21: 115. 1897.
Hybrid formula: Calanthe tankervilleae (Banks) M. W. Chase, Christenh. & Schuit. × [Calanthe × veitchii Paxton & Lindl.].
Type: Not designated.
Calanthe × inspirata (Veitch) Z. H. Feng, comb. nov. — Basionym: × Phaiocalanthe inspirata Veitch, Gard. Chron. ser. 3, 22: 315. 1897.
Hybrid formula: Calanthe masuca (D. Don) Lindl. × Calanthe tankervilleae (Banks) M. W. Chase, Christenh. & Schuit.
Type: Not designated.
Poaceae
Anthoxanthum × zinserlingii (Tzvelev) Z. H. Feng, comb. nov. — Basionym: Hierochloe × zinserlingii Tzvelev, Novosti Sist. Vyssh. Rast. 10: 83. 1973.
Hybrid formula: Anthoxanthum monticola (Bigelow) Veldkamp × Anthoxanthum nitens (Weber) Y. Schouten & Veldkamp.
Type: RUSSIA. Ural. on the cliffs of Mount Sablja, 20 July 1926, G. Zinserling 504 (holotype: LE [barcode] LE 01078250, digital image!).
× Pseudelymus versicolor (A. P. Khokhr.) Z. H. Feng, comb. nov. — Basionym: Elymus × versicolor A. P. Khokhr., Biol. Rast. Fl. Sev. Dal'n. Vostok.: 13. 1981.
Hybrid formula: Elymus macrourus (Turcz. ex Steud.) Tzvelev × Pseudoroegneria gmelinii (Trin. ex Schrad.) Sennikov.
Type: RUSSIA. Magadan Region. Srednekansky District, mountain Zamkovyi, mosses by the rock, 22 August 1979, S. Ershova, P. Khokryakov, A. P. Khokryakov s. n. (holotype: MHA [barcode] MHA 0359393, digital image!; isotypes: MAG [barcodes] MAG 0000235-MAG 0000241, digital images!; VLA [barcode] VLA 00001831).
Rosaceae
× Arsorbus abscondita (Kovanda) Z. H. Feng, comb. nov. — Basionym: Sorbus × abscondita Kovanda, Verh. Zool.-Bot. Ges. Österreich 133: 339. 1996.
Hybrid formula: Aria danubialis (Jáv.) Sennikov & Kurtto × Sorbus aucuparia L.
Type: Czech Republic. Bohemia occid.-centr.: in angulo clivi Pochvalovská stráň dicti situ septentr.-orient. a pago Pochvalov (distr. Louny); solo calcifero-schistaceo, alt. 490 m, 25 May 1992, M. Kovanda s. n. (holotype: WU; isotypes: PR).
Eriolobus angustifolius (Aiton) Su Liu ex Z. H. Feng, comb. nov. — Basionym: Pyrus angustifolia Aiton, Hort. Kew. [W. Aiton] 2: 176. 1789.
Neotype (designated by Li et al.
: 71): USA. North Carolina, locality unknown, s. d.,
Greene s. n. (neotype: K [barcode] K 000758465, digital image!).
Eriolobus coronarius (L.) Su Liu ex Z. H. Feng, comb. nov. — Basionym: Pyrus coronaria L., Sp. Pl. [Linnaeus] 1: 480. 1753.
Type: Provenance not given. s. d., Linnaeus s. n., Herb. Linn. No. 647.5 (holotype: LINN [barcode] LINN 647.5, digital image!).
Eriolobus ioensis (Alph. Wood) Su Liu ex Z. H. Feng, comb. & stat. nov. — Basionym: Pyrus coronaria var. ioensis Alph. Wood, Class-book Bot. (ed. 1861): 333. 1861.
Neotype (designated by Langenfeld
[132] | Langenfeld V. T. Apple Trees: Morphological Evolution, Phylogeny, Geography and Systematics of the Genus. Zinatne Publishing, Riga; 1991, pp. 232. |
[132]
: 153): USA. Missouri, Daviess, near Pattonsburg, thickets, limestone uplands, 14 June 1924,
E. J. Palmer 25466 (neotype: S).
× Eriomalus Su Liu ex Z. H. Feng, nothogen. nov.
Hybrid formula: Eriolobus (DC.) M. Roem. × Malus Mill.
× Eriomalus heterophylla (Spach) Z. H. Feng, comb. nov. — Basionym: Malus × heterophylla Spach, Hist. Nat. Vég. (Spach) 2: 138. 1834, pro sp.
Hybrid formula: Eriolobus coronarius (L.) Su Liu ex Z. H. Feng × Malus domestica (Suckow) Borkh.
Type: Not designated.
× Eriomalus soulardii (L. H. Bailey) Su Liu ex Z. H. Feng, comb. nov. — Basionym: Pyrus × soulardii L. H. Bailey (‘soulardi’), Amer. Garden 12: 472. 1891, pro sp.
Hybrid formula: Eriolobus ioensis (Alph. Wood) Su Liu ex Z. H. Feng × Malus domestica (Suckow) Borkh.
Type: Not designated.
× Tormariosorbus liljeforsii (T. C. G. Rich) Z. H. Feng, comb. nov. — Basionym: Sorbus × liljeforsii T. C. G. Rich, Nordic J. Bot. 25(5-6): 339. 2008.
Hybrid formula: Sorbus aucuparia L. × [× Tormariosorbus intermedia (Ehrh.) J. M. H. Shaw].
Type: Sweden. Västergötland: Finnerödja, Brinken, 10 July 1920, J. A. O. Skårman s. n. (holotype: UPS).
Saxifragaceae
Micranthes × crawfordii (E. S. Marshall) Z. H. Feng, comb. nov. — Basionym: Saxifraga × crawfordii E. S. Marshall, J. Bot. 47: 98. 1938.
Hybrid formula: Micranthes nivalis (L.) Small × Micranthes stellaris (L.) Galasso, Banfi & Soldano.
Type: Not designated.
Thelypteridaceae
× Abacopterella jerdonii (Ching) Z. H. Feng, comb. nov. — Basionym: Cyclosorus jerdonii Ching, Bull. Fan Mem. Inst. Biol. Bot. 8: 228. 1938.
Hybrid formula: Abacopteris nudata (Roxb.) S. E. Fawc. & A. R. Sm. × Christella procera (D. Don) Mazumdar.
Type: SIKKIM [India]. Jerdon s. n. (holotype: K [barcode] K 000951517, digital image!; isotypes: B [barcodes] B 200041322, B 200041323, B 200041324, digital images!).
× Chririx Z. H. Feng, nothogen. nov.
Hybrid formula: Christella H. Lév. × Grypothrix (Holttum) S. E. Fawc. & A. R. Sm.
× Chririx gogoii (Fraser-Jenk.) Z. H. Feng, comb. nov. — Basionym: Thelypteris × gogoii Fraser-Jenk., New Sp. Syndr. Indian Pteridol.: 248. 1997.
Hybrid formula: Christella procera (D. Don) Mazumdar × Grypothrix triphylla (Sw.) S. E. Fawc. & A. R. Sm.
Type: INDIA. Assam, Jorhat District, Goiaghat, shortly north of Kosarihat village, 21 December 1995, C. R Fraser-Jenkins 23846, FN 3001 (holotype: BM).
× Chririx thwaitesii (Hook.) Z. H. Feng, comb. nov. — Basionym: Meniscium × thwaitesii Hook., Fil. Exot. 2: t. 83. 1858.
Hybrid formula: Christella parasitica (L.) H. Lév. × Grypothrix triphylla (Sw.) S. E. Fawc. & A. R. Sm.
Type: SRI LANKA. s. d., G. H. K. Thwaites 3145 (holotype: K [barcode] K 000951504, digital image!; isotypes: B [barcodes] B 200068843, B 200068848, BM [barcode] BM 001044868, GH [barcode] GH 00112616, K [barcode] K 000951505, digital image!, L [barcode] L 0061084, P [barcodes] P 00643850, P 00643851, UC [barcode] UC 267428).
× Chrismatopteris varievenulosa (Viane) Z. H. Feng, comb. nov. — Basionym: Thelypteris × varievenulosa Viane, Bull. Soc. Roy. Bot. Belgique 118(1): 49. 1985.
Hybrid formula: Christella hispidula (Decne.) Holttum × Pneumatopteris afra (Christ) Holttum.
Type: WEST AFRICA [IVORY COAST]. Mt. Tonkoui. Wet, shaded trackside at ca. 900 m alt., among Cyathea camerooniana and Thelypteris afra, 29 July 1979, R. Viane 1030 (holotype: GENT [barcode] GENT 0000090035227, digital image!; isotype: GENT [barcode] GENT 0000090035166, digital image!).
× Christelliopsis nepalensis (Fraser-Jenk.) Z. H. Feng, comb. nov. — Basionym: Thelypteris × nepalensis Fraser-Jenk., Taxon. Revis. Indian Subcontinental Pteridophytes: 188. 2008.
Hybrid formula: Christella dentata (Forssk.) Brownsey & Jermy × Menisciopsis penangiana (Hook.) S. E. Fawc. & A. R. Sm.
Type: NEPAL. Nawalparasi District, Lumbini Zone, c. 1 km northeast of Daunne near top of pass over Siwalik ridge, shortly south of Dumkibas, east of Butwal on road to Narayanghat, 28 November 2005, C. R. Fraser-Jenkins 31892 (holotype: TAIF [barcode] TAIF 309339, digital image!).
× Glaphyronisciopsis Z. H. Feng, nothogen. nov.
Hybrid formula: Glaphyropteridopsis Ching × Menisciopsis (Holttum) S. E. Fawc. & A. R. Sm.
× Glaphyronisciopsis occulta (C. Hope) Z. H. Feng, comb. nov. — Basionym: Nephrodium occultum C. Hope, J. Bombay Nat. Hist. Soc. 12: 627, t. 13. 1899.
Hybrid formula: Glaphyropteridopsis erubescens (Wall. ex Hook.) Ching × Menisciopsis penangiana (Hook.) S. E. Fawc. & A. R. Sm.
Lectotype (designated by Singh and Rawat
[133] | Singh, R. K., Rawat, V. K. New combinations for eleven nothospecies in Thelypteridaceae. J. Biodivers. Conservation 2024, 8(4), 26-41. |
[133]
: 40): INDIA. North-western India, Uttarakhand, 1890,
P. W. Mackinnon & V. A. Mackinnon s. n. (lectotype: BM [barcode] BM 001044870, digital image!).
Grypothrix × pseudoliukiuensis (Seriz.) Z. H. Feng, comb. nov. — Basionym: Thelypteris × pseudoliukiuensis Seriz., J. Phytogeogr. Taxon. 29(1): 25. 1981.
Hybrid formula: Grypothrix cuspidata (Blume) S. E. Fawc. & A. R. Sm. × Grypothrix triphylla (Sw.) S. E. Fawc. & A. R. Sm.
Type: RYUKYU ISLAND. Mt. Komidake, alt. ca. 90 m, 9 August 1971, S. Serizawa 14678 (holotype: AICH).